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Hansen's spike-moss

Eaton's spike-moss

Habit Plants terrestrial, forming loose to clustered mats. Plants terrestrial, forming tiny (1–4 cm), dense clumps.
Stems

not readily fragmenting, prostrate, upperside and underside structurally different, irregularly forked, branches determinate, tips upturned.

short-creeping, unbranched or few-forked, flat, not articulate, glabrous.

Leaves

with underside leaves slightly longer and narrower than upperside leaves, otherwise monomorphic, not clearly ranked, tightly appressed, ascending, green or green with red spots, or reddish, linear-lanceolate (underside) to linear-triangular (upperside), (2–)3–4.5 × 0.5–0.6 mm;

abaxial ridges present;

base abruptly adnate, pubescent (sometimes glabrous);

margins ciliate, cilia white to white opaque, strongly appressed and ascending, 0.03–0.1 mm;

apex with bristle white to white-opaque, 0.5–1.4 mm (those on underside leaves sometimes 1/4–1/2 longer than those on upperside leaves).

delicate, papery.

Strobili

solitary, 5–7 mm;

sporophylls ovate-deltate to ovate-triangular, abaxial ridges not prominent, base glabrous, margins short-ciliate, apex bristled.

solitary, 2–3 mm;

sporophylls ovate-lanceolate, strongly keeled toward tip, keel dentate, base glabrous, margins serrate, apex long-acuminate.

Rhizophores

borne on upperside of stems, throughout stem length, 0.25–0.45 mm diam.

axillary, 0.02–0.04 mm diam.

Lateral

leaves spreading, well spaced or crowded toward stem tip, green, ovate to ovate-oblong, 1–1.5 × 0.5–0.9 mm;

base rounded;

margins transparent, serrate;

apex acute.

Median

leaves lanceolate, 0.8–1.2 × 0.3–0.35 mm;

base oblique;

margins transparent, serrate;

apex bristled;

bristle to 1/3 length of leaf.

Selaginella hansenii

Selaginella eatonii

Habitat Cliffs and rocky slopes or on igneous rock Hammocks and sink holes in limestone soil
Elevation 330–1350 m (1100–4400 ft) 0 m (0 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; West Indies in the Bahamas
[BONAP county map]
Discussion

Leaf dimorphism in Selaginella hansenii is only slightly and inconsistently expressed; the upperside leaves tend to be more lanceolate, short, and slightly thick, whereas the underside leaves tend to be more linear, longer, and thinner, but in some specimens the leaves are monomorphic. Red leaves are rare within Selaginella subg. Tetragonostachys, otherwise found in the flora only occasionally in S. rupestris. Such leaves are more common in S. steyermarkii Alston from southern Mexico and Guatemala and S. sartorii Hieronymus from Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Selaginella eatonii is a minute species, easy to distinguish by its long-bristled leaf apex, iridescent leaf surface, and somewhat transparent sporophylls on the underside of the stem. It does not have any close relatives among the species in the flora. Selaginella eatonii may best be placed within subg. Heterostachys, with which it shares flattened strobili, keeled sporophylls, and a partial laminar flap on the sporophylls. Selaginella eatonii does not have strongly dimorphic sporophylls or a very well-defined laminar flap, and therefore I prefer to treat it here with the other heterophyllous species of subg. Stachygynandrum until a detailed study of subg. Heterostachys can be made.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Selaginellaceae > Selaginella > subg. Tetragonostachys Selaginellaceae > Selaginella > subg. Stachygynandrum
Sibling taxa
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eclipes, S. eremophila, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
Synonyms Diplostachyum eatonii
Name authority Hieronymus: Hedwigia 39: 301. (1900) Hieronymus ex Small: Ferns Trop. Florida 67. (1918)
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