Selaginella eremophila |
Selaginella douglasii |
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desert spike-moss |
Douglas selaginella, Douglas' spike-moss, Douglas' spikemoss clubmoss, lesser clubmoss |
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Habit | Plants on rock or terrestrial, forming dense mats. | Plants on rock or terrestrial, forming loose mats. |
Stems | not readily fragmenting, prostrate, upperside and underside structurally different, irregularly forked; branches determinate, tips upturned. |
long-creeping, branched, branches 2–3-forked, flat, not articulate, glabrous. |
Leaves | conspicuously dimorphic, in 8 ranks, tightly appressed, ascending, green; abaxial ridges present; apex with deciduous, twisted, transparent bristle ± 0.3 mm, becoming acute to slightly mucronate in oldest branches. |
delicate and papery. |
Strobili | solitary, 3–8 mm; sporophylls ovate-deltate, abaxial ridges not prominent, base glabrous, margins ciliate, apex acute to mucronate. |
paired, 0.6–1.1 cm; sporophylls monomorphic, ovate-lanceolate, keeled, keel not dentate, base glabrous, margins green, entire or with a few scattered, short cilia, apex acute to acuminate. |
Rhizophores | borne on upperside of stems, throughout stem length, 0.2 mm diam. |
borne on underside of stems throughout stem length or restricted to proximal ± 2/3 of main stem or axillary throughout stem, 0.2–0.4 mm diam. |
Underside | leaves lanceolate to lanceolate-elliptic (on central ranks) or falcate (on marginal ranks), 2–2.7 × 0.5–0.7 mm; base decurrent, glabrous; margins ciliate, cilia transparent to opaque, spreading, 0.04–0.1 mm. |
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Upperside | leaves lanceolate, 1.3–1.4 × 0.3–0.4 mm; base abruptly adnate, pubescent, hairs often running along groove; margins ciliate, cilia transparent to opaque, spreading, ca. 0.1 mm. |
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Lateral | leaves spreading or slightly ascending, distant, shiny green becoming shiny brown, with orange or red spot or entirely reddish, ovate to ovate-oblong or oblong, 1.5–3.2 X (1–)1.5–2.2 mm; base auriculate, basiscopic auricle conspicuous, acroscopic auricle inconspicuous or base ± rounded; margins green, ciliate toward auricles, otherwise entire; apex rounded to obtuse or truncate. |
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Median | leaves ovate-oblong, (1.8–)2–2.2 × 1–1.3 mm; base auriculate, outer auricle larger than inner one; margins green, ciliate at auricles, otherwise entire; apex abruptly cuspidate to bristled. |
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Selaginella eremophila |
Selaginella douglasii |
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Habitat | Rocky and sandy slopes, in open rock or crevices or in soil | Rocky slopes, mossy rock, rock crevices, in partial shade, often along river banks |
Elevation | 130–1000 m (400–3300 ft) | 100–800 m (300–2600 ft) |
Distribution |
AZ; CA; Mexico in Baja California
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ID; OR; WA
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Discussion | Selaginella eremophila is most closely related to the Mexican S. parishii L. Underwood and S. landii Greenman & Pfeiffer. In S. eremophila and the following two species, S. arizonica and S. peruviana, the leaves are arranged in 8 conspicuous ranks: 3 underside (2 marginal, 1 central), 2 lateral, and 3 upperside (2 marginal, 1 central). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Selaginella douglasii, with no close relatives in the flora, is easy to identify by its shiny green leaves when young, turning shiny light brown when old, with an orange to red spot at the base, or totally reddish. Its closest relative is the Mexican S. delicatissima Linden ex A. Braun. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Selaginellaceae > Selaginella > subg. Tetragonostachys | Selaginellaceae > Selaginella > subg. Stachygynandrum |
Sibling taxa | ||
Synonyms | Lycopodium douglasii | |
Name authority | Maxon: Smithsonian Misc. Collect. 72: 3–5. (1920) | (Hooker & Greville) Spring: Bull. Acad. Roy. Sci. Bruxelles 10: 138. (1843) |
Web links |