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buck's meadow spike-moss, hidden spike-moss

desert spike-moss

Habit Plants terrestrial, forming loose to dense mats. Plants on rock or terrestrial, forming dense mats.
Stems

short-creeping, branched, branches 1–2-forked, flat, not articulate, glabrous.

not readily fragmenting, prostrate, upperside and underside structurally different, irregularly forked;

branches determinate, tips upturned.

Leaves

papery, delicate.

conspicuously dimorphic, in 8 ranks, tightly appressed, ascending, green;

abaxial ridges present;

apex with deciduous, twisted, transparent bristle ± 0.3 mm, becoming acute to slightly mucronate in oldest branches.

Strobili

solitary or paired, lax, flattened, 1–4 cm;

sporophylls ovate to ovate-deltate, strongly keeled, keel dentate, base glabrous, slightly cordate to rounded, margins serrate, apex acuminate;

megasporophylls larger and wider than microsporophylls, usually on underside of strobili.

solitary, 3–8 mm;

sporophylls ovate-deltate, abaxial ridges not prominent, base glabrous, margins ciliate, apex acute to mucronate.

Rhizophores

throughout stem length, 0.06–0.1 mm diam.

borne on upperside of stems, throughout stem length, 0.2 mm diam.

Lateral

leaves nearly perpendicular to stem, green, ovate to ovate-elliptic, 1–2 × 0.5–1.3 mm;

base rounded to slightly subcordate;

margins slightly transparent, serrate;

apex acute.

Median

leaves ovate to ovate-lanceolate or lanceolate, 1–1.8 × 0.4–0.8 mm;

base rounded to oblique;

margins green, serrate;

apex abruptly tapered, long-acuminate to bristled, frequently transparent, midrib extending into apex.

Underside

leaves lanceolate to lanceolate-elliptic (on central ranks) or falcate (on marginal ranks), 2–2.7 × 0.5–0.7 mm;

base decurrent, glabrous;

margins ciliate, cilia transparent to opaque, spreading, 0.04–0.1 mm.

Upperside

leaves lanceolate, 1.3–1.4 × 0.3–0.4 mm;

base abruptly adnate, pubescent, hairs often running along groove;

margins ciliate, cilia transparent to opaque, spreading, ca. 0.1 mm.

Selaginella eclipes

Selaginella eremophila

Habitat Moist to wet, calcareous habitats, swamps, meadows, pastures, open woods, or rarely on rock Rocky and sandy slopes, in open rock or crevices or in soil
Elevation 0–100 m (0–300 ft) 130–1000 m (400–3300 ft)
Distribution
from FNA
AR; IA; IL; IN; MI; MO; NY; OK; WI; ON; QC
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from FNA
AZ; CA; Mexico in Baja California
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Discussion

Selaginella eclipes, a member of the S. apoda complex, may prove to be better treated as a subspecies of S. apoda (W. R. Buck 1977). It is recognized here at the specific level to highlight the problems within this species complex. Further research is needed to elucidate the relationships among the species of the complex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Selaginella eremophila is most closely related to the Mexican S. parishii L. Underwood and S. landii Greenman & Pfeiffer. In S. eremophila and the following two species, S. arizonica and S. peruviana, the leaves are arranged in 8 conspicuous ranks: 3 underside (2 marginal, 1 central), 2 lateral, and 3 upperside (2 marginal, 1 central).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Selaginellaceae > Selaginella > subg. Stachygynandrum Selaginellaceae > Selaginella > subg. Tetragonostachys
Sibling taxa
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eclipes, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
Name authority W. R. Buck: Canad. J. Bot. 55: 366. (1977) Maxon: Smithsonian Misc. Collect. 72: 3–5. (1920)
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