Selaginella douglasii |
Selaginella weatherbiana |
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Douglas selaginella, Douglas' spike-moss, Douglas' spikemoss clubmoss, lesser clubmoss |
Weatherby's spike-moss |
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Habit | Plants on rock or terrestrial, forming loose mats. | Plants on rock, forming clumps. |
Stems | long-creeping, branched, branches 2–3-forked, flat, not articulate, glabrous. |
radially symmetric, underground (rhizomatous) and aerial, not readily fragmenting, irregularly forked; rhizomatous and aerial stems often with 1 branch arrested, budlike, tips straight; aerial stems erect, less often ascending, cespitose, stout, branches not conspicuously arrested, budlike branches mostly near base. |
Leaves | delicate and papery. |
dimorphic, not clearly ranked. |
Strobili | paired, 0.6–1.1 cm; sporophylls monomorphic, ovate-lanceolate, keeled, keel not dentate, base glabrous, margins green, entire or with a few scattered, short cilia, apex acute to acuminate. |
solitary, (0.7–)1–3 cm; sporophylls narrowly ovate-lanceolate to lanceolate, abaxial ridges prominent, base glabrous, margins denticulate to short-ciliate, apex keeled, bristled. |
Rhizophores | borne on underside of stems throughout stem length or restricted to proximal ± 2/3 of main stem or axillary throughout stem, 0.2–0.4 mm diam. |
borne on upperside of stems, mostly restricted to rhizomatous stems or to lower 1/2 of aerial stems, 0.16–0.26(–3) mm diam. |
Lateral | leaves spreading or slightly ascending, distant, shiny green becoming shiny brown, with orange or red spot or entirely reddish, ovate to ovate-oblong or oblong, 1.5–3.2 X (1–)1.5–2.2 mm; base auriculate, basiscopic auricle conspicuous, acroscopic auricle inconspicuous or base ± rounded; margins green, ciliate toward auricles, otherwise entire; apex rounded to obtuse or truncate. |
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Median | leaves ovate-oblong, (1.8–)2–2.2 × 1–1.3 mm; base auriculate, outer auricle larger than inner one; margins green, ciliate at auricles, otherwise entire; apex abruptly cuspidate to bristled. |
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Rhizomatous | stem leaves persistent, loosely appressed, ascending, often incurved, scalelike. |
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Aerial | stem leaves tightly appressed, ascending, green, linear-lanceolate to narrowly lanceolate, 1.7–2.4 × 0.36–0.43 mm; abaxial ridges prominent; base cuneate and decurrent on main stem or rounded and abruptly adnate on apical branch portions, glabrous or pubescent; margins short-ciliate at base, cilia transparent, spreading, denticulate, and ascending toward apex, 0.03–0.06 mm; apex keeled; bristle transparent to opaque or yellowish to brownish (on old leaves), puberulent to smooth, 0.3–0.6(–0.7) mm. |
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2n | = 18. |
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Selaginella douglasii |
Selaginella weatherbiana |
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Habitat | Rocky slopes, mossy rock, rock crevices, in partial shade, often along river banks | Exposed or shaded granitic rock outcrops, ledges, cliffs, or in rock crevices |
Elevation | 100–800 m (300–2600 ft) | 1600–3000 m (5200–9800 ft) |
Distribution |
ID; OR; WA
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CO; NM |
Discussion | Selaginella douglasii, with no close relatives in the flora, is easy to identify by its shiny green leaves when young, turning shiny light brown when old, with an orange to red spot at the base, or totally reddish. Its closest relative is the Mexican S. delicatissima Linden ex A. Braun. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
One of the most striking features of Selaginella weatherbiana is that at branch forks the larger branch continues to grow as a vegetative shoot, and the smaller one usually forms a strobilus. Therefore, the strobili appear to be lateral rather than terminal. Selaginella weatherbiana grows in close association with S. underwoodii (R. M. Tryon 1955). The two species (as well as S. mutica var. mutica) are very often mixed on herbarium specimens. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Selaginellaceae > Selaginella > subg. Stachygynandrum | Selaginellaceae > Selaginella > subg. Tetragonostachys |
Sibling taxa | ||
Synonyms | Lycopodium douglasii | |
Name authority | (Hooker & Greville) Spring: Bull. Acad. Roy. Sci. Bruxelles 10: 138. (1843) | R. M. Tryon: Amer. Fern J. 40: 69. (1950) |
Web links |