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Douglas selaginella, Douglas' spike-moss, Douglas' spikemoss clubmoss, lesser clubmoss

Wallace's selaginella, Wallace's spike-moss

Habit Plants on rock or terrestrial, forming loose mats. Plants on rock or terrestrial, forming loose or compact mats.
Stems

long-creeping, branched, branches 2–3-forked, flat, not articulate, glabrous.

radially symmetric, creeping or decumbent, not readily fragmenting, irregularly forked, without budlike arrested branches, tips straight;

main stem long, indeterminate, lateral branches determinate, ascending, 1–2-forked.

Leaves

delicate and papery.

monomorphic, in ± alternate pseudowhorls of 4, tightly or loosely appressed, ascending, green, linear-lanceolate, (1.5–)1.8–3.5 × 0.39–0.66 mm;

abaxial ridges well defined;

base rounded and adnate or cuneate and slightly decurrent on fleshy, loosely appressed stem leaves (from wet places), pubescent, seldom glabrous;

margins short-ciliate to denticulate, cilia transparent, spreading at base, dentiform, and ascending toward apex, 0.03–0.06(–0.1) mm;

apex keeled and obtuse, sometimes attenuate or plane and attenuate, abruptly short- to long-bristled;

bristle transparent to whitish, puberulent, sometimes breaking off, (0.16–)0.2–0.46(–0.9) mm.

Strobili

paired, 0.6–1.1 cm;

sporophylls monomorphic, ovate-lanceolate, keeled, keel not dentate, base glabrous, margins green, entire or with a few scattered, short cilia, apex acute to acuminate.

often paired, 1–4.5(–9) cm;

sporophylls deltate-ovate (mostly on exposed and compact mats) or lanceolate-ovate (on loose, spreading mats from wet places), abaxial ridges well defined, base glabrous, margins short-ciliate to denticulate, apex keeled, abruptly short-bristled, seldom tapering into bristle.

Rhizophores

borne on underside of stems throughout stem length or restricted to proximal ± 2/3 of main stem or axillary throughout stem, 0.2–0.4 mm diam.

borne on upperside of stems, throughout stem length, 0.23–0.36(–0.4) mm diam.

Lateral

leaves spreading or slightly ascending, distant, shiny green becoming shiny brown, with orange or red spot or entirely reddish, ovate to ovate-oblong or oblong, 1.5–3.2 X (1–)1.5–2.2 mm;

base auriculate, basiscopic auricle conspicuous, acroscopic auricle inconspicuous or base ± rounded;

margins green, ciliate toward auricles, otherwise entire;

apex rounded to obtuse or truncate.

Median

leaves ovate-oblong, (1.8–)2–2.2 × 1–1.3 mm;

base auriculate, outer auricle larger than inner one;

margins green, ciliate at auricles, otherwise entire;

apex abruptly cuspidate to bristled.

Selaginella douglasii

Selaginella wallacei

Habitat Rocky slopes, mossy rock, rock crevices, in partial shade, often along river banks On dry, exposed cliffs, rocky slopes, rocky knolls, or sandy-gravelly soil or on moist, shaded, rocky banks or in meadows
Elevation 100–800 m (300–2600 ft) 0–2000 m (0–6600 ft)
Distribution
from FNA
ID; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Selaginella douglasii, with no close relatives in the flora, is easy to identify by its shiny green leaves when young, turning shiny light brown when old, with an orange to red spot at the base, or totally reddish. Its closest relative is the Mexican S. delicatissima Linden ex A. Braun.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Selaginella wallacei is extremely variable depending on its habitat (R. M. Tryon 1955). Plants in dry, exposed conditions have short stems, form compact mats with tightly appressed leaves adnate to the stem, and have a rather keeled, abruptly bristled apex. Plants from moist habitats have long stems, form rather moderately long-creeping mats, and have less appressed, decurrent, fleshy leaves, with a more plane-attenuate apex that gradually tapers into a bristle. Plants from exposed, dry conditions sometimes are confused with S. scopulorum, but they have a keeled apex with well-defined ridges on the abaxial groove whereas in S. scopulorum the leaf apex is ± plane and attenuate, and the ridges on the abaxial groove are not prominent. Plants from moist habitats somewhat resemble plants of S. underwoodii.

R. M. Tryon (1955) found strobili 9 cm long in Selaginella wallacei, the longest strobili known within subg. Tetragonostachys and comparable only to those of S. oregana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Selaginellaceae > Selaginella > subg. Stachygynandrum Selaginellaceae > Selaginella > subg. Tetragonostachys
Sibling taxa
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
Synonyms Lycopodium douglasii
Name authority (Hooker & Greville) Spring: Bull. Acad. Roy. Sci. Bruxelles 10: 138. (1843) Hieronymus: Hedwigia 39: 297. (1900)
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