Selaginella douglasii |
Selaginella sibirica |
|
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Douglas selaginella, Douglas' spike-moss, Douglas' spikemoss clubmoss, lesser clubmoss |
northern selaginella, Siberian spike-moss |
|
Habit | Plants on rock or terrestrial, forming loose mats. | Plants on rock or terrestrial, forming discrete long-spreading mats or seldom cushionlike mats. |
Stems | long-creeping, branched, branches 2–3-forked, flat, not articulate, glabrous. |
radially symmetric, creeping or decumbent, not readily fragmenting, irregularly forked, without budlike arrested branches, tips straight; main stem indeterminate, lateral branches conspicuously or inconspicuously determinate, often strongly ascending, 1–3-forked. |
Leaves | delicate and papery. |
monomorphic, in alternate pseudowhorls of 5, tightly appressed, ascending, green, linear-lanceolate to narrowly lanceolate, 2–3.5 × 0.35–0.5 mm (smaller on lateral branches); abaxial ridges prominent; base cuneate and decurrent to rounded and adnate on young lateral branches or buds, glabrous or sometimes pubescent; margins long-ciliate, cilia transparent, spreading to ascending, 0.07–0.17 mm; apex keeled, truncate in profile, obtuse to attenuate; bristle white to whitish or transparent, puberulent, 0.45–0.8 mm. |
Strobili | paired, 0.6–1.1 cm; sporophylls monomorphic, ovate-lanceolate, keeled, keel not dentate, base glabrous, margins green, entire or with a few scattered, short cilia, apex acute to acuminate. |
solitary, 0.5–2.5 cm; sporophylls deltate-ovate to ovate-lanceolate, abaxial ridges well defined, base glabrous, margins ciliate, apex truncate in profile, bristled. |
Rhizophores | borne on underside of stems throughout stem length or restricted to proximal ± 2/3 of main stem or axillary throughout stem, 0.2–0.4 mm diam. |
borne on upperside of stems, throughout stem length, 0.2–0.37 mm diam. |
Lateral | leaves spreading or slightly ascending, distant, shiny green becoming shiny brown, with orange or red spot or entirely reddish, ovate to ovate-oblong or oblong, 1.5–3.2 X (1–)1.5–2.2 mm; base auriculate, basiscopic auricle conspicuous, acroscopic auricle inconspicuous or base ± rounded; margins green, ciliate toward auricles, otherwise entire; apex rounded to obtuse or truncate. |
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Median | leaves ovate-oblong, (1.8–)2–2.2 × 1–1.3 mm; base auriculate, outer auricle larger than inner one; margins green, ciliate at auricles, otherwise entire; apex abruptly cuspidate to bristled. |
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2n | = 18. |
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Selaginella douglasii |
Selaginella sibirica |
|
Habitat | Rocky slopes, mossy rock, rock crevices, in partial shade, often along river banks | Dry, alpine, rocky slopes, rock crevices, granite rock, limestone boulders, sandstone, bare open grassy tundra |
Elevation | 100–800 m (300–2600 ft) | 130–2400 m (400–7900 ft) |
Distribution |
ID; OR; WA
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AK; NT; YT; Asia in Japan and the former Soviet republics |
Discussion | Selaginella douglasii, with no close relatives in the flora, is easy to identify by its shiny green leaves when young, turning shiny light brown when old, with an orange to red spot at the base, or totally reddish. Its closest relative is the Mexican S. delicatissima Linden ex A. Braun. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Selaginella sibirica is most closely allied to S. rupestris. In addition to differences noted in the descriptions, it can be distinguished from S. rupestris by the numerous marginal cilia on the leaves and by the transparent sporophyll margins; S. rupestris has a variable number (usually few) of marginal cilia and nontransparent sporophyll margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Selaginellaceae > Selaginella > subg. Stachygynandrum | Selaginellaceae > Selaginella > subg. Tetragonostachys |
Sibling taxa | ||
Synonyms | Lycopodium douglasii | S. rupestris |
Name authority | (Hooker & Greville) Spring: Bull. Acad. Roy. Sci. Bruxelles 10: 138. (1843) | (J. Milde) Hieronymus: Hedwigia 39: 290. (1900) |
Web links |