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Douglas selaginella, Douglas' spike-moss, Douglas' spikemoss clubmoss, lesser clubmoss

northern selaginella, Siberian spike-moss

Habit Plants on rock or terrestrial, forming loose mats. Plants on rock or terrestrial, forming discrete long-spreading mats or seldom cushionlike mats.
Stems

long-creeping, branched, branches 2–3-forked, flat, not articulate, glabrous.

radially symmetric, creeping or decumbent, not readily fragmenting, irregularly forked, without budlike arrested branches, tips straight;

main stem indeterminate, lateral branches conspicuously or inconspicuously determinate, often strongly ascending, 1–3-forked.

Leaves

delicate and papery.

monomorphic, in alternate pseudowhorls of 5, tightly appressed, ascending, green, linear-lanceolate to narrowly lanceolate, 2–3.5 × 0.35–0.5 mm (smaller on lateral branches);

abaxial ridges prominent;

base cuneate and decurrent to rounded and adnate on young lateral branches or buds, glabrous or sometimes pubescent;

margins long-ciliate, cilia transparent, spreading to ascending, 0.07–0.17 mm;

apex keeled, truncate in profile, obtuse to attenuate;

bristle white to whitish or transparent, puberulent, 0.45–0.8 mm.

Strobili

paired, 0.6–1.1 cm;

sporophylls monomorphic, ovate-lanceolate, keeled, keel not dentate, base glabrous, margins green, entire or with a few scattered, short cilia, apex acute to acuminate.

solitary, 0.5–2.5 cm;

sporophylls deltate-ovate to ovate-lanceolate, abaxial ridges well defined, base glabrous, margins ciliate, apex truncate in profile, bristled.

Rhizophores

borne on underside of stems throughout stem length or restricted to proximal ± 2/3 of main stem or axillary throughout stem, 0.2–0.4 mm diam.

borne on upperside of stems, throughout stem length, 0.2–0.37 mm diam.

Lateral

leaves spreading or slightly ascending, distant, shiny green becoming shiny brown, with orange or red spot or entirely reddish, ovate to ovate-oblong or oblong, 1.5–3.2 X (1–)1.5–2.2 mm;

base auriculate, basiscopic auricle conspicuous, acroscopic auricle inconspicuous or base ± rounded;

margins green, ciliate toward auricles, otherwise entire;

apex rounded to obtuse or truncate.

Median

leaves ovate-oblong, (1.8–)2–2.2 × 1–1.3 mm;

base auriculate, outer auricle larger than inner one;

margins green, ciliate at auricles, otherwise entire;

apex abruptly cuspidate to bristled.

2n

= 18.

Selaginella douglasii

Selaginella sibirica

Habitat Rocky slopes, mossy rock, rock crevices, in partial shade, often along river banks Dry, alpine, rocky slopes, rock crevices, granite rock, limestone boulders, sandstone, bare open grassy tundra
Elevation 100–800 m (300–2600 ft) 130–2400 m (400–7900 ft)
Distribution
from FNA
ID; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; YT; Asia in Japan and the former Soviet republics
[BONAP county map]
Discussion

Selaginella douglasii, with no close relatives in the flora, is easy to identify by its shiny green leaves when young, turning shiny light brown when old, with an orange to red spot at the base, or totally reddish. Its closest relative is the Mexican S. delicatissima Linden ex A. Braun.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Selaginella sibirica is most closely allied to S. rupestris. In addition to differences noted in the descriptions, it can be distinguished from S. rupestris by the numerous marginal cilia on the leaves and by the transparent sporophyll margins; S. rupestris has a variable number (usually few) of marginal cilia and nontransparent sporophyll margins.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Selaginellaceae > Selaginella > subg. Stachygynandrum Selaginellaceae > Selaginella > subg. Tetragonostachys
Sibling taxa
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
Synonyms Lycopodium douglasii S. rupestris
Name authority (Hooker & Greville) Spring: Bull. Acad. Roy. Sci. Bruxelles 10: 138. (1843) (J. Milde) Hieronymus: Hedwigia 39: 290. (1900)
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