Selaginella douglasii |
Selaginella rupincola |
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Douglas selaginella, Douglas' spike-moss, Douglas' spikemoss clubmoss, lesser clubmoss |
rockloving spikemoss |
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Habit | Plants on rock or terrestrial, forming loose mats. | Plants on rock or terrestrial, forming loose clumps. |
Stems | long-creeping, branched, branches 2–3-forked, flat, not articulate, glabrous. |
radially symmetric, underground (rhizomatous) and aerial, not readily fragmenting, irregularly forked; both rhizomatous and aerial stems often with 1 branch arrested, budlike, tips straight; rhizomatous stems hard to distinguish on wholly creeping plants; aerial stems erect or ascending, sometimes decumbent to slightly creeping, budlike arrested branches restricted mostly near stem base. |
Leaves | delicate and papery. |
dimorphic, not clearly ranked. |
Strobili | paired, 0.6–1.1 cm; sporophylls monomorphic, ovate-lanceolate, keeled, keel not dentate, base glabrous, margins green, entire or with a few scattered, short cilia, apex acute to acuminate. |
solitary, 0.5–2.5(–3.5) cm; sporophylls lanceolate, strongly tapering toward tip, abaxial ridges prominent, base glabrous, margins short-ciliate, apex long-bristled. |
Rhizophores | borne on underside of stems throughout stem length or restricted to proximal ± 2/3 of main stem or axillary throughout stem, 0.2–0.4 mm diam. |
borne on upperside of stems, restricted to lower stems or throughout stem length, 0.3–0.5 mm diam. |
Lateral | leaves spreading or slightly ascending, distant, shiny green becoming shiny brown, with orange or red spot or entirely reddish, ovate to ovate-oblong or oblong, 1.5–3.2 X (1–)1.5–2.2 mm; base auriculate, basiscopic auricle conspicuous, acroscopic auricle inconspicuous or base ± rounded; margins green, ciliate toward auricles, otherwise entire; apex rounded to obtuse or truncate. |
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Median | leaves ovate-oblong, (1.8–)2–2.2 × 1–1.3 mm; base auriculate, outer auricle larger than inner one; margins green, ciliate at auricles, otherwise entire; apex abruptly cuspidate to bristled. |
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Rhizomatous | stem leaves persistent or deciduous, tightly appressed, scalelike. |
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Aerial | stem leaves appressed, ascending, green, linear-lanceolate, 3–4.7 × 0.45–0.65 mm; abaxial ridges present; base abruptly adnate, rounded, pubescent; margins long-ciliate, cilia white to whitish, spreading, 0.1–0.2 mm; apex not keeled to slightly keeled; bristle white to whitish or yellowish to greenish near base, puberulent, 0.65–1.85 mm (1/3–1/2 length of leaves). |
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Selaginella douglasii |
Selaginella rupincola |
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Habitat | Rocky slopes, mossy rock, rock crevices, in partial shade, often along river banks | Exposed ledges and rock, steep slopes, rock crevices or gravelly soil |
Elevation | 100–800 m (300–2600 ft) | 1000–2000 m (3300–6600 ft) |
Distribution |
ID; OR; WA
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AZ; NM; TX; Mexico
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Discussion | Selaginella douglasii, with no close relatives in the flora, is easy to identify by its shiny green leaves when young, turning shiny light brown when old, with an orange to red spot at the base, or totally reddish. Its closest relative is the Mexican S. delicatissima Linden ex A. Braun. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Selaginella rupincola is allied to S. bigelovii. It is one of the presumed parents of S. × neomexicana (see discussion). In addition to characteristics given, it can be separated from S. bigelovii in having hairs often running along the ridges of the abaxial groove, whereas S. bigelovii has nonhairy ridges on the abaxial groove. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Selaginellaceae > Selaginella > subg. Stachygynandrum | Selaginellaceae > Selaginella > subg. Tetragonostachys |
Sibling taxa | ||
Synonyms | Lycopodium douglasii | |
Name authority | (Hooker & Greville) Spring: Bull. Acad. Roy. Sci. Bruxelles 10: 138. (1843) | L. Underwood: Bull. Torrey Bot. Club 25: 129. (1898) |
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