Selaginella douglasii |
Selaginella ludoviciana |
|
---|---|---|
Douglas selaginella, Douglas' spike-moss, Douglas' spikemoss clubmoss, lesser clubmoss |
gulf spike-moss, Louisiana spike-moss |
|
Habit | Plants on rock or terrestrial, forming loose mats. | Plants terrestrial, forming diffuse mats. |
Stems | long-creeping, branched, branches 2–3-forked, flat, not articulate, glabrous. |
long-creeping, usually ascending, sparsely branched, branches mostly simple or 1-forked, flat, not articulate, glabrous. |
Leaves | delicate and papery. |
delicate, papery. |
Strobili | paired, 0.6–1.1 cm; sporophylls monomorphic, ovate-lanceolate, keeled, keel not dentate, base glabrous, margins green, entire or with a few scattered, short cilia, apex acute to acuminate. |
solitary or paired, lax, flattened, 0.4–0.7(–1.5) cm; sporophylls very strongly keeled, keel dentate, base slightly cordate to rounded, margins transparent, sparsely serrate, apex acuminate; megasporophylls larger and wider than microsporophylls, usually on underside of strobili. |
Rhizophores | borne on underside of stems throughout stem length or restricted to proximal ± 2/3 of main stem or axillary throughout stem, 0.2–0.4 mm diam. |
axillary, 0.1–0.2 mm diam. |
Lateral | leaves spreading or slightly ascending, distant, shiny green becoming shiny brown, with orange or red spot or entirely reddish, ovate to ovate-oblong or oblong, 1.5–3.2 X (1–)1.5–2.2 mm; base auriculate, basiscopic auricle conspicuous, acroscopic auricle inconspicuous or base ± rounded; margins green, ciliate toward auricles, otherwise entire; apex rounded to obtuse or truncate. |
leaves well spaced, green, ovate to ovate-lanceolate, 1.6–2.65 × 0.98–1.64 mm; base slightly cordate; margins transparent, serrate; apex acute to slightly obtuse, conspicuously ending in teeth. |
Median | leaves ovate-oblong, (1.8–)2–2.2 × 1–1.3 mm; base auriculate, outer auricle larger than inner one; margins green, ciliate at auricles, otherwise entire; apex abruptly cuspidate to bristled. |
leaves spaced, ovate-lanceolate to narrowly ovate-lanceolate (on basal stems), 1.3–2 × 0.4–0.8 mm; base oblique on inner side, rounded and prominent on outer side; margins transparent, serrate; apex long-acuminate to bristled. |
Selaginella douglasii |
Selaginella ludoviciana |
|
Habitat | Rocky slopes, mossy rock, rock crevices, in partial shade, often along river banks | Swamps, stream banks, ditch banks, or moist ravines of calcareous ledges |
Elevation | 100–800 m (300–2600 ft) | 0–50 m (0–200 ft) |
Distribution |
ID; OR; WA
|
AL; FL; GA; LA; MS |
Discussion | Selaginella douglasii, with no close relatives in the flora, is easy to identify by its shiny green leaves when young, turning shiny light brown when old, with an orange to red spot at the base, or totally reddish. Its closest relative is the Mexican S. delicatissima Linden ex A. Braun. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Among the species in the flora, Selaginella ludoviciana is most closely related to S. apoda (see discussion) and has often been included in S. apoda. W. R. Buck and T. W. Lucansky (1976) concluded that two species should be recognized based on anatomic and morphologic data. A close examination of distribution of sporangia in the strobili in many specimens reveals that sporangial arrangement may be more variable than reported by H. T. Horner Jr. and H. J. Arnott (1963). All species in the S. apoda complex (see S. apoda and S. eclipes for discussion on the complex) have sporophylls with fused blade tissue. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Selaginellaceae > Selaginella > subg. Stachygynandrum | Selaginellaceae > Selaginella > subg. Stachygynandrum |
Sibling taxa | ||
Synonyms | Lycopodium douglasii | Lycopodium ludovicianum, Diplostachyum ludovicianum |
Name authority | (Hooker & Greville) Spring: Bull. Acad. Roy. Sci. Bruxelles 10: 138. (1843) | (A. Braun) A. Braun: Ann. Sci. Nat., Bot. sér. 4, 13: 58. (1860) |
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