Selaginella apoda |
Selaginella scopulorum |
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meadow spike-moss, sélaginelle apode |
cliff selaginella, cliff spikemoss, compact spikemoss, Rocky Mountain spike-moss |
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Habit | Plants terrestrial, forming loose or clustered mats. | Plants terrestrial or on rock, forming cushionlike or rather loose mats. |
Stems | prostrate to short-creeping, sparsely branched, branches mostly simple or 1-forked, flat, not articulate, glabrous. |
decumbent or creeping, not readily fragmenting, irregularly forked, without budlike arrested branches, tips straight; main stem upperside and underside structurally slightly different, conspicuously or inconspicuously indeterminate, lateral branches radially symmetric, conspicuously determinate, strongly ascending, 1–2-forked. |
Leaves | delicate, papery. |
monomorphic, in poorly defined pseudowhorls of 4 or 6, tightly appressed, ascending, green, linear-lanceolate to linear, in lateral ranks sometimes falcate, 2.5–4(–4.3) X 0.5–0.75 mm (upperside leaves smaller than underside, smaller also on ascending buds); abaxial ridges present; base (on main stem) decurrent, oblique, and glabrous on underside leaves, slightly decurrent to adnate, oblique, and glabrous or rarely puberulent on upperside leaves; margins usually short-ciliate, cilia transparent, spreading or ascending at base, denticulate and ascending on distal 2/3, 0.02–0.07(–0.15) mm; apex plane or sometimes slightly keeled, obtuse to attenuate, abruptly bristled; bristle whitish, transparent to opaque, with few teeth or smooth, 0.5–1.1 mm. |
Strobili | paired or solitary, lax, flattened, 1–2 cm; sporophylls ovate to ovate-deltate, strongly keeled, keel dentate, base slightly cordate to rounded, margins with scattered teeth, apex acuminate; megasporophylls larger and wider than microsporophylls, usually on underside of strobili. |
solitary, (0.5–)1–3(–4.5) cm; sporophylls ovate-lanceolate, lanceolate, or seldom ovate, usually tapering toward apex, abaxial ridges well defined, base glabrous, margins proximally short-ciliate to denticulate, lacking cilia apically, apex usually attenuate or slightly keeled, short-bristled. |
Rhizophores | axillary, throughout stem length or restricted to proximal 1/3 of stem, 0.05–0.1 mm diam. |
borne on upperside of stems, throughout stem length, 0.25–0.45 mm diam. |
Lateral | leaves distant, green, ovate to ovate-lanceolate, 1.35–2.25 × 0.75–1.35 mm; base slightly cordate; margins green or with 1 row of transparent cells, serrate; apex acute, ending in teeth. |
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Median | leaves ovate-lanceolate, 1–1.6 × 0.45–0.7 mm; base oblique on inner side, rounded and prominent on outer side; margins green or with row of transparent cells, serrate; apex straight, acuminate to long-acuminate. |
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Selaginella apoda |
Selaginella scopulorum |
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Habitat | Swamps, meadows, marshes, pastures, damp lawns, open woods, and stream banks, in basic to acidic soil | Rocky alpine tundra, subalpine meadows, dry cliffs, rocky slopes, rock crevices, granitic outcrops and ledges, sandstone outcrops, in soil pockets among rocks, or sandy or granitic soil |
Elevation | 0–100 m (0–300 ft) | 700–3700 m (2300–12100 ft) |
Distribution |
AL; AR; CT; DE; FL; GA; IL; IN; KY; LA; MA; MD; ME; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WV; Mexico in Chihuahua; s to Chiapas
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AZ; CA; CO; ID; MT; NM; OR; UT; WA; WY; AB; BC
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Discussion | Selaginella apoda is the central component of a taxonomically difficult species complex of eastern North America. It is closely related to S. eclipes (see discussion) and S. ludoviciana. Naturally occurring and experimental hybrids between S. apoda and S. ludoviciana have been reported (P. Somers and W. R. Buck 1975; T. R. Webster 1990). Also, some evidence indicates that hybrids may occur between S. apoda and S. eclipes. More studies are needed in this complex. The species in the S. apoda complex may be best classified under subg. Homostachys of J. G. Baker (1883, 1887), with which they share flattened strobili and larger sporophylls (megasporophylls) that are usually in the same plane as the vegetative lateral leaves. They are, however, treated here with the other heterophyllous species of subg. Stachygynandrum until a reassessment of the classification of the genus Selaginella can be made. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Selaginella scopulorum is a member of the S. densa complex, in which there is a clear need for more systematic studies. Some specimens of S. scopulorum from Montana, Wyoming, and Colorado have more conspicuous whitish bristles than those elsewhere and are difficult to distinguish from S. densa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Selaginellaceae > Selaginella > subg. Stachygynandrum | Selaginellaceae > Selaginella > subg. Tetragonostachys |
Sibling taxa | ||
Synonyms | Lycopodium apodum, Diplostachyum apodum | S. densa var. scopulorum, S. engelmannii var. scopulorum |
Name authority | (Linnaeus) C. Morren: in Martius et al., Fl. Bras. 1(2): 119. (1840) | Maxon: Amer. Fern J. 11: 36. (1921) |
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