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broad-leaf stonecrop, Pacific stonecrop, spatula-leaf stonecrop

Havard's stonecrop

Habit Herbs, perennial, mat-forming, glabrous. Herbs or subshrubs, perennial, not tufted, glabrous.
Stems

rhizomatous, procumbent or creeping, much-branched, bearing terminal rosettes.

procumbent, creeping, or spreading, with ascending or erect branches, (tuberculate), not bearing rosettes.

Flowering shoots

erect, simple, 3–14 cm;

leaf blades spatulate-oblong or elliptic-oblong, base not spurred;

offsets not formed.

erect, simple, 1–5 cm, (papillose);

leaf blades elliptic, base short-spurred;

offsets not formed.

Leaves

alternate, spreading, petiolate;

blade green, often glaucous or pruinose, spatulate, terete to laminar, 7–19 × 4.5–10 mm, base not spurred, not scarious, apex rounded or truncate, submucronate, (surfaces papillose marginally).

alternate, (imbricate to subimbricate), erect to spreading, sessile;

blade bright green, sometimes glaucous, suboblong to ovate, somewhat flattened to terete, 4–9 × 1–2 mm, base widened, short-spurred, not scarious, apex obtuse, (surfaces smooth or papillose).

Inflorescences

cymes, ca. 30-flowered, ca. 3-branched;

branches not recurved, forked;

bracts oblong-spatulate or linear, ca. 3 cm, base not spurred.

3-parted cymes, 2–10-flowered or flowers solitary, simple or monochasially 1-branched;

branches erect to slightly recurved, not forked;

bracts similar to leaves, sometimes imbricate.

Pedicels

2–8 mm.

absent or to 1.5 mm.

Flowers

5-merous;

sepals spreading to erect, connate basally, green or yellow-green, glaucous or pruinose, lanceolate, linear-lanceolate, oblong-ovate, or obovate, equal, ca. 2.5 × 1.5 mm, apex acute or obtuse;

petals widely spreading from short, erect base, distinct or slightly connate basally, yellow, linear to oblanceolate, not carinate, 4.5–9 mm, apex acute;

filaments yellow;

anthers yellow;

nectar scales yellow, reniform or nearly square.

5-merous;

sepals erect to suberect, distinct basally, green to pinkish, linear or lanceolate, slightly unequal, 3–5 × 0.2–1 mm, apex obtuse;

petals spreading, distinct basally, white, suboblong, not carinate, 5–6.5 mm, apex obtuse, shortly mucronate;

filaments white to pale pink;

anthers red or purplish;

nectar scales whitish to pale pink, oblong, (retuse).

Carpels

divergent in fruit, connate basally, brown.

divergent or stellately spreading in fruit, distinct, red to purple or stramineous with reddish to purplish striations.

2n

= 30.

= ca. 36–50, 68–72.

Sedum spathulifolium

Sedum havardii

Phenology Flowering spring–fall.
Habitat Igneous rock outcrops or talus in oak-pinyon woodlands and chaparral
Elevation 1500-2500 m (4900-8200 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The mature carpels of Sedum spathulifolium have five ribs and prominent lips along the adaxial suture. The flowers are sweetly fragrant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Primary rosettes 2.5-3.7 cm diam.; leaf blades not pruinose, glaucous or not, 1.4-2.1 mm thick; flowers 1.2-1.6 cm diam.
var. spathulifolium
1. Primary rosettes 1.5-2.1 cm diam.; leaf blades pruinose, not glaucous, 2.1-2.5 mm thick; flowers 1-1.2 cm diam.
var. pruinosum
Source FNA vol. 8, p. 222. FNA vol. 8, p. 214.
Parent taxa Crassulaceae > Sedum Crassulaceae > Sedum
Sibling taxa
S. acre, S. albomarginatum, S. album, S. annuum, S. borschii, S. cockerellii, S. debile, S. divergens, S. glaucophyllum, S. havardii, S. hispanicum, S. lanceolatum, S. laxum, S. leibergii, S. lineare, S. mexicanum, S. moranii, S. nanifolium, S. nevii, S. niveum, S. nuttallii, S. oblanceolatum, S. obtusatum, S. ochroleucum, S. oreganum, S. oregonense, S. praealtum, S. pulchellum, S. pusillum, S. radiatum, S. robertsianum, S. rupestre, S. rupicola, S. sarmentosum, S. sexangulare, S. stelliforme, S. stenopetalum, S. ternatum, S. villosum, S. wrightii
S. acre, S. albomarginatum, S. album, S. annuum, S. borschii, S. cockerellii, S. debile, S. divergens, S. glaucophyllum, S. hispanicum, S. lanceolatum, S. laxum, S. leibergii, S. lineare, S. mexicanum, S. moranii, S. nanifolium, S. nevii, S. niveum, S. nuttallii, S. oblanceolatum, S. obtusatum, S. ochroleucum, S. oreganum, S. oregonense, S. praealtum, S. pulchellum, S. pusillum, S. radiatum, S. robertsianum, S. rupestre, S. rupicola, S. sarmentosum, S. sexangulare, S. spathulifolium, S. stelliforme, S. stenopetalum, S. ternatum, S. villosum, S. wrightii
Subordinate taxa
S. spathulifolium var. pruinosum, S. spathulifolium var. spathulifolium
Name authority Hooker: Fl. Bor.-Amer. 1: 227. (1832) Rose: in N. L. Britton et al., N. Amer. Fl. 22: 74. 1905 (as havardi),
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