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broad-leaf stonecrop, Pacific stonecrop, spatula-leaf stonecrop

orpin, stonecrop

Habit Herbs, perennial, mat-forming, glabrous. Herbs, annual, biennial, or perennial, rarely subshrubs, not viviparous, 0.2–10 dm, glabrous or hairy.
Stems

rhizomatous, procumbent or creeping, much-branched, bearing terminal rosettes.

erect, ascending, procumbent, or creeping, usually much-branched, sometimes succulent, usually hardly secondarily thickened, base sometimes slightly woody.

Flowering shoots

erect, simple, 3–14 cm;

leaf blades spatulate-oblong or elliptic-oblong, base not spurred;

offsets not formed.

Leaves

alternate, spreading, petiolate;

blade green, often glaucous or pruinose, spatulate, terete to laminar, 7–19 × 4.5–10 mm, base not spurred, not scarious, apex rounded or truncate, submucronate, (surfaces papillose marginally).

deciduous or persistent, rarely rosulate, sometimes forming dense rosettes, usually alternate, sometimes opposite or in whorls of 3–5[–6], ± alike, petiolate or sessile, not connate basally;

blade linear to orbiculate or spatulate, terete, semiterete, or subterete, rarely laminar, 0.1–8 cm, usually ± succulent, base spurred or not, margins entire, (with 1 abaxial apical hydathode);

veins not conspicuous.

Inflorescences

cymes, ca. 30-flowered, ca. 3-branched;

branches not recurved, forked;

bracts oblong-spatulate or linear, ca. 3 cm, base not spurred.

terminal or axillary cymes (sometimes compound, cincinnate, less often simple).

Pedicels

2–8 mm.

present or absent.

Flowers

5-merous;

sepals spreading to erect, connate basally, green or yellow-green, glaucous or pruinose, lanceolate, linear-lanceolate, oblong-ovate, or obovate, equal, ca. 2.5 × 1.5 mm, apex acute or obtuse;

petals widely spreading from short, erect base, distinct or slightly connate basally, yellow, linear to oblanceolate, not carinate, 4.5–9 mm, apex acute;

filaments yellow;

anthers yellow;

nectar scales yellow, reniform or nearly square.

erect, (3–)5–8[–12]-merous;

sepals distinct and spurred or slightly connate basally, (green), equal or strongly unequal;

petals spreading or erect at anthesis, usually distinct or slightly connate basally, [connate for 1/3–2/3 their lengths], yellow, white, pink, purple, or reddish, sometimes red-lineolate (often with reddish keel and adaxial subapical appendage), margins entire;

calyx and corolla not circumscissile;

nectaries oblong, square, transversely oblong, reniform, or spatulate;

stamens 2 times as many as sepals, rarely as many as sepals;

filaments of epipetalous stamens usually adnate basally, filaments of antipetalous stamens usually distinct, rarely connate basally;

pistils erect, slightly connate basally to almost distinct, (usually sessile with broad base);

styles equaling or shorter than ovary.

Fruits

erect or spreading.

Carpels

divergent in fruit, connate basally, brown.

Seeds

ovoid to ellipsoid, (usually ca. 1 mm), reticulate or reticulate-papillose.

2n

= 30.

= 4–ca. 640.

Sedum spathulifolium

Sedum

Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; Mexico; Central America; Europe; Asia; n Africa; e Africa; Atlantic Islands (Iceland); Indian Ocean Islands (Madagascar)
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The mature carpels of Sedum spathulifolium have five ribs and prominent lips along the adaxial suture. The flowers are sweetly fragrant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Amerosedum Á. Löve & D. Löve; Breitungia Á. Löve & D. Löve; Chetyson Rafinesque; Clausenella Á. Löve & D. Löve; Cockerellia (R. T. Clausen & C. H. Uhl) Á. Löve & D. Löve; Hjaltalinia Á. Löve & D. Löve

Species 420–470 (41 in the flora).

Many genera of Crassulaceae currently recognized are derived from Sedum, the largest genus in the family. Molecular studies by H. ’t Hart (1995), R. C. H. J. van Ham and ’t Hart (1998), ’t Hart et al. (1999), S. Mayuzumi and H. Ohba (2004), and M. E. Mort et al. (2001) have formed the basis for recognizing segregate genera when the nomenclatural, biogeographic, and morphological data also have been available. Such data are still lacking for many other groups, and these are retained in Sedum pending further study. R. T. Clausen (1975) provided the most comprehensive treatment to date of native and naturalized Sedum in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Primary rosettes 2.5-3.7 cm diam.; leaf blades not pruinose, glaucous or not, 1.4-2.1 mm thick; flowers 1.2-1.6 cm diam.
var. spathulifolium
1. Primary rosettes 1.5-2.1 cm diam.; leaf blades pruinose, not glaucous, 2.1-2.5 mm thick; flowers 1-1.2 cm diam.
var. pruinosum
1. Annual or biennial herbs
→ 2
1. Perennial herbs or subshrubs
→ 14
2. Biennial herbs
→ 3
2. Annual herbs
→ 7
3. Flowering shoots forming axillary offsets
→ 4
3. Flowering shoots not forming axillary offsets
→ 5
4. Filaments white; carpels stellately spreading in fruit
S. radiatum
4. Filaments yellow; carpels divergent in fruit.
S. stenopetalum
5. Petals canary yellow; sepals slightly connate basally, glabrous.
S. leibergii
5. Petals pink-red or purple to white; sepals distinct, glabrous or glandular-hairy
→ 6
6. Stems and leaves glabrous; sepals glabrous; petals white to purple, 4-10 mm.
S. pulchellum
6. Stems and leaves hirtellous and glandular-hairy; sepals glandular-hairy; petals pink-red, 3-4.5 mm
S. villosum
7. Stems glandular-hairy (at least distally) or hirtellous
→ 8
7. Stems glabrous
→ 9
8. Stems hirtellous; leaf blades hirtellous and densely glandular-hairy; sepals 2-4 mm.
S. villosum
8. Stems glandular-hairy; leaf blades usually glabrous, rarely glandular-hairy; sepals ca. 2 mm.
S. hispanicum
9. Petals 4-11 mm
→ 10
9. Petals 1.4-5 mm
→ 12
10. Flowering shoots without axillary offsets; c, e United States.
S. pulchellum
10. Flowering shoots with axillary offsets; w United States
→ 11
11. Carpels stellately spreading in fruit.
S. radiatum
11. Carpels divergent in fruit.
S. stenopetalum
12. Petals white, ovate-elliptic.
S. pusillum
12. Petals yellow, elliptic-oblong or narrowly lanceolate
→ 13
13. Leaf blade bases short-spurred; petals elliptic-oblong; c United States.
S. nuttallii
13. Leaf blade bases not spurred; petals narrowly lanceolate; Greenland.
S. annuum
14. Petals white, creamy white, yellowish white, pale yellow, yellowish, white tinged with purple or streaked with pink, pinkish white, greenish white, purple, pink-red, or pink
→ 15
14. Petals yellow, yellowish green, golden yellow, pale orange, or pale orange suffused with pink
→ 32
15. Flowers mostly 4-merous, sometimes 5-6(-7)-merous
→ 16
15. Flowers mostly 5(-8)-merous
→ 19
16. Leaf blades of flowering stems linear, base sagittately spurred.
S. pulchellum
16. Leaf blades of flowering shoots obovate to obovate-spatulate or elliptic, narrowly elliptic, oblong or linear, base short-spurred
→ 17
17. Leaves whorled, usually 3, sometimes 2 or 4 per whorl, sometimes decussate.
S. ternatum
17. Leaves alternate
→ 18
18. Leaf blades not glaucous; petals acute apically; primary rosettes usually 1 cm diam.
S. nevii
18. Leaf blades usually glaucous; petals acuminate apically; primary rosettes usually 1-2.7 cm diam.
S. glaucophyllum
19. Flowers pungent, musky scented (unique in the flora area); petals sharply curved outward 2/3 from erect base, distal 1/3 divergent
S. wrightii
19. Flowers without pungent scent; petals erect, widely spreading from base, or spreading from middle, not sharply curved outward
→ 20
20. Inflorescences panicles or paniculate cymes
→ 21
20. Inflorescences 1-3(-5)-parted or branched cymes, corymbs, or corymbiform cymes
→ 25
21. Stems creeping and short-ascending, basally densely glandular- pubescent; petals 2-4.5 mm.
S. album
21. Stems rootstocks or stout rhizomes, bearing rosettes, glabrous; petals 4-11.5 mm
→ 22
22. Leaf blades narrowly obovate, strongly pruinose, not glaucous; sepals 4-7 mm.
S. oblanceolatum
22. Leaf blades obovate, spatulate, or oblanceolate, not strongly pruinose, sometimes glaucous; sepals 1.8-6 mm
→ 23
23. Petals 4-10 mm; sepals loosely appressed to corolla, 1.8-5.3 mm; flowering shoots 2-10(-12) cm.
S. obtusatum
23. Petals ca. (4-)7-11.5 mm; sepals closely appressed to corolla, (2-)2.6-5.1 mm; flowering shoots (6-)10-28(-40) cm
→ 24
24. Petals pink, pinkish white, or white, apices obtuse; sepals 2.6-5.1 mm
S. laxum
24. Petals creamy white, yellowish white, or pale yellow, apices abruptly pointed; sepals 3.2-3.5 mm.
S. oregonense
25. Leaf blades of flowering shoots linear or linear-lanceolate to linear-oblanceolate
→ 26
25. Leaf blades of flowering shoots lanceolate, obovate to oblanceolate, oblanceolate-elliptic, elliptic-ovate, oblong-spatulate, oblanceolate-oblong, spatulate, or elliptic to elliptic-oblong
→ 29
26. Offsets often produced from axils of leaves and bracts.
S. stenopetalum
26. Offsets not formed from axils of leaves and bracts
→ 27
27. Sepals connate basally.
S. ochroleucum
27. Sepals distinct
→ 28
28. Sepals lanceolate or ovate-elliptic.
S. villosum
28. Sepals linear to narrowly oblong.
S. stelliforme
29. Carpels divergent or stellately spreading in fruit.
S. havardii
29. Carpels erect in fruit
→ 30
30. Inflorescences (4-)10-27-flowered cymes; pedicels 1-3.5 mm; leaf blades 9.5-15 mm
S. cockerellii
30. Inflorescences 2-9-flowered cymes or solitary flowers; pedicels absent or to 1 mm, or pedicels 3-8 mm; leaf blades 3-8 mm
→ 31
31. Pedicels absent or to 1 mm.
S. niveum
31. Pedicels 3-8 mm.
S. villosum
32. Leaves whorled or opposite and decussate (rarely alternate)
→ 33
32. Leaves alternate
→ 37
33. Leaves usually opposite and decussate (rarely alternate)
→ 34
33. Leaves whorled
→ 35
34. Petals usually distinct, sometimes connate basally; carpels widely spreading in fruit; leaf blades green or reddish green, not glaucous, suborbiculate to spatulate-obovate or oblong.
S. divergens
34. Petals connate basally; carpels erect or ascending in fruit; leaf blades pale green speckled with pink, sometimes pink or red, glaucous, elliptic, ovate-elliptic, oblanceolate, or obovate.
S. debile
35. Leaves 4(-5) per whorl, blade linear-elliptic or linear-lanceolate; flowering shoots erect.
S. mexicanum
35. Leaves 3(-4) per whorl, blade narrowly rhombic-elliptic to widely lanceolate or linear-lanceolate to linear; flowering shoots ascending, pendulous, or creeping
→ 36
36. Leaf blades narrowly rhombic-elliptic to widely lanceolate; flowering shoots creeping or ascending.
S. sarmentosum
36. Leaf blades linear-lanceolate to linear; flowering shoots ascending or pendulous.
S. lineare
37. Inflorescences paniculate cymes
→ 38
37. Inflorescences corymbs, corymbiform cymes, or cymes, sometimes subscorpioid
→ 42
38. Flowering shoots glandular-pubescent.
S. moranii
38. Flowering shoots glabrous
→ 39
39. Subshrubs; leaf blades elliptic-oblanceolate, 40-80 × 13-25 mm; petals widely spreading.
S. praealtum
39. Perennial herbs; leaf blades obovate, oblanceolate, oblanceolate-oblong, or spatulate, (4-)12-28(-67) × 4-11(-18) mm; petals erect proximally, spreading or divergent distally
→ 40
40. Leaf blade margins white; 300-900 m.
S. albomarginatum
40. Leaf blade margins not white; 300-3700 m
→ 41
41. Flowering shoots 2-10(-12) cm.
S. obtusatum
41. Flowering shoots (6-)10-28 cm.
S. oregonense
42. Leaf blades linear or linear-lanceolate to oblong
→ 43
42. Leaf blades spatulate, orbiculate to broadly ovate, ovate, obovate, lanceolate, elliptic, elliptic-lanceolate, elliptic-oblong, cuneate, or triangular-ovate
→ 46
43. Petals 3-4 mm; leaves usually in 6 rows.
S. sexangulare
43. Petals 5-10 mm; leaves on vegetative shoots spirally arranged or alternate, not in rows
→ 44
44. Flowering shoots with axillary offsets; leaf blade bases scarious.
S. stenopetalum
44. Flowering shoots without axillary offsets; leaf blade bases not scarious
→ 45
45. Leaf blades linear-lanceolate; sepals lanceolate, 3-6 mm, densely glandular-pubescent; petals erect or suberect, lanceolate, 8-10 mm.
S. ochroleucum
45. Leaf blades linear to oblong; sepals ovate, 2-3(-3.3) mm, glabrous or sparsely glandular-pubescent; petals spreading, oblong, 6-7 mm.
S. rupestre
46. Plants not forming rosettes or offsets
→ 47
46. Plants forming rosettes, or offsets formed at time of flowering
→ 49
47. Plants tufted.
S. robertsianum
47. Plants mat-forming
→ 48
48. Leaf blades orbiculate to broadly ovate.
S. nanifolium
48. Leaf blades triangular-ovate.
S. acre
49. Leaf blades obovate, lanceolate, elliptic, elliptic-lanceolate, elliptic-ovate, ovate, elliptic-oblong, or linear
→ 50
49. Leaf blades spatulate, oblong-spatulate, or cuneate
→ 53
50. Carpels erect in fruit
→ 51
50. Carpels divergent in fruit
→ 52
51. Leaves not easily detached; petals canary yellow to golden yellow.
S. lanceolatum
51. Leaves easily detached; petals deep yellow.
S. rupicola
52. Leaf blades linear to elliptic-oblong.
S. stenopetalum
52. Leaf blades usually obovate or elliptic, rarely lanceolate.
S. borschii
53. Flowering shoots glandular-pubescent.
S. moranii
53. Flowering shoots glabrous
→ 54
54. Petals widely spreading from short, erect base; filaments yellow.
S. spathulifolium
54. Petals basally erect, divergent distally; filaments white, greenish, greenish white, pink, or yellowish
→ 55
55. Petals not carinate, pink, pinkish white, or white to yellowish white; leaf blades 10-50 × 6-33 mm, base not spurred.
S. laxum
55. Petals carinate, yellow; leaf blades 7-12 × 4.4-9 mm, base short-spurred.
S. oreganum
Source FNA vol. 8, p. 222. FNA vol. 8, p. 199. Author: Hideaki Ohba.
Parent taxa Crassulaceae > Sedum Crassulaceae
Sibling taxa
S. acre, S. albomarginatum, S. album, S. annuum, S. borschii, S. cockerellii, S. debile, S. divergens, S. glaucophyllum, S. havardii, S. hispanicum, S. lanceolatum, S. laxum, S. leibergii, S. lineare, S. mexicanum, S. moranii, S. nanifolium, S. nevii, S. niveum, S. nuttallii, S. oblanceolatum, S. obtusatum, S. ochroleucum, S. oreganum, S. oregonense, S. praealtum, S. pulchellum, S. pusillum, S. radiatum, S. robertsianum, S. rupestre, S. rupicola, S. sarmentosum, S. sexangulare, S. stelliforme, S. stenopetalum, S. ternatum, S. villosum, S. wrightii
Subordinate taxa
S. spathulifolium var. pruinosum, S. spathulifolium var. spathulifolium
S. acre, S. albomarginatum, S. album, S. annuum, S. borschii, S. cockerellii, S. debile, S. divergens, S. glaucophyllum, S. havardii, S. hispanicum, S. lanceolatum, S. laxum, S. leibergii, S. lineare, S. mexicanum, S. moranii, S. nanifolium, S. nevii, S. niveum, S. nuttallii, S. oblanceolatum, S. obtusatum, S. ochroleucum, S. oreganum, S. oregonense, S. praealtum, S. pulchellum, S. pusillum, S. radiatum, S. robertsianum, S. rupestre, S. rupicola, S. sarmentosum, S. sexangulare, S. spathulifolium, S. stelliforme, S. stenopetalum, S. ternatum, S. villosum, S. wrightii
Name authority Hooker: Fl. Bor.-Amer. 1: 227. (1832) Linnaeus: Sp. Pl. 1: 430. (1753): Gen. Pl. ed. 5, 197. 1754 ,
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