Sclerocactus parviflorus |
Sclerocactus papyracanthus |
|
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devil's-claw cactus, small flower fishhook cactus |
grama-grass cactus, paper-spine cactus, paperspine fishhook cactus, toumeya |
|
Stems | unbranched or branched near base, depressed-spheric, spheric, cylindric, or elongate-cylindric, 4.5–45 × 3.5–14.5 cm; ribs (10–)13(–16), tubercles evident on ribs. |
unbranched, cylindric or obconic cylindric, 2–7.5(–8) × 1.2–2.5 cm; ribs not evident, tubercles prominent. |
Spines | frequently obscuring stems; radial spines 8–17 per areole, usually white, sometimes brown or purplish pink, 6–36 mm; central spines (3–)4–6(–8) per areole, 1–5 hooked; abaxial central spine usually purple black, hooked, 15–72 × 0.6–1 mm; lateral spines 2–6 per areole, similar to abaxial, usually shorter, usually not hooked; adaxial central spine usually white (rarely gray, straw colored, pink, or reddish brown), flat to angled or terete; straight (rarely contorted), 17–82(–90) × 0.7–1.5(–3) mm. |
dense, obscuring stems; radial spines 5–10 per areole, white, straight, flat, (2–)3–5 × 0.3–0.6 mm; central spines 1(–4) per areole; abaxial central spine 1 per areole, whitish to tan or gray, straight, wavy, twisting, or curling, flat, flexible, papery, with obscure adaxial midrib, sometimes pointing upward and obscuring apex of plant, lacking hook, 15–45(–50) × 1–1.5 mm; lateral or adaxial central spines 0(–3) per areole, white to tan, (3–)20–40 × 0.5–1 mm. |
Flowers | funnelform to campanulate, (2–)3–5.7(–7) × 2.5–5.5(–8) cm; outer tepals with greenish or brownish purple midstripes and rose to purple, pink, yellow, or white margins, oblanceolate, 25–50 × 6–10 mm, mucronate, margins entire; inner tepals rose to purple, pink, or yellow (rarely white), oblanceolate, 15–55 × 6–12 mm, margins entire to fringed, apex mucronate; filaments purple, yellow, or green; anthers yellow; ovary papillate, appearing granular. |
funnelform to narrowly campanulate, 2–2.5 × 1–2.5 cm; outer tepals with green-purple to red-brown midstripes and cream to white margins, cuneate-spatulate, usually 9–20 × 1–3 mm, finely toothed; inner tepals white with brown midstripes, largest tepals oblanceolate, 15–20 × 3.5–4.5 mm, apex acute to mucronate; filaments white to greenish yellow; anthers cream to pale yellow. |
Fruits | irregularly dehiscent, green turning reddish pink, 10–30 × 10–15 mm; scales few, membranous fringed. |
indehiscent or irregularly dehiscent, green, subspheric, 4–6 × 3–5 mm, dry at maturity; scales few or none. |
Seeds | dark brown to black, 2.5–3.5 × 1.5–3 mm; testa with rounded papillae. |
black, 2.5–3 × 2–2.5 mm, shiny; testa with fine, rounded papillae. |
Sclerocactus parviflorus |
Sclerocactus papyracanthus |
|
Phenology | Flowering late Apr–May. | Flowering Apr–Jun. |
Habitat | Sandy, gravelly, or clay hills, mesas, and washes, desert grasslands or saltbush, sagebrush, rabbitbrush, and blackbrush communities, pinyon-juniper woodlands | Desert grasslands, pinyon-juniper woodlands, Chihuahuan desert scrub |
Elevation | 1000-2100 m (3300-6900 ft) | 1500-2200 m (4900-7200 ft) |
Distribution |
AZ; CO; NM; UT
|
AZ; NM; TX |
Discussion | Sclerocactus parviflorus has an extensive range and great morphologic variation. Many named segregates have been proposed (L. D. Benson 1982; K. D. Heil and J. M. Porter 1994; F. Hochstätter 1995b) and some of these clearly merit recognition at the subspecific or varietal level. However, we are not attempting to subdivide this very complex group. Considerable work remains in understanding morphologic diversification within this species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
With long, flexuous, flattened spines and pale flowers, Sclerocactus papyracanthus is surprisingly cryptic in the field. There has been a long debate concerning its taxonomic placement. Chloroplast DNA analyses of J. M. Porter et al. (2000) unambiguously placed this species within Sclerocactus, a position first suggested based on morphologic evidence (K. D. Heil et al. 1981; C. Glass and R. A. Foster 1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 202. | FNA vol. 4, p. 205. |
Parent taxa | Cactaceae > subfam. Cactoideae > Sclerocactus | Cactaceae > subfam. Cactoideae > Sclerocactus |
Sibling taxa | ||
Synonyms | Echinocactus parviflorus, Echinocact parviflorus var. havasupaiensis, Echinocact parviflorus var. roseus, Ferocactus parviflorus, Pediocactus parviflorus, S. contortus, S. havasupaiensis var. roseus, S. intermedius, S. parviflorus subsp. havasupaiensis, S. parviflorus var. intermedius, S. parviflorus subsp. terrae-canyonae, S. terrae-canyonae, S. whipplei var. intermedius, S. whipplei var. roseus | Mammillaria papyracantha, Pediocactus papyracanthus, Toumeya papyracantha |
Name authority | Clover & Jotter: Bull. Torrey Bot. Club 68: 419, fig. 8. (1941) | (Engelmann) N. P. Taylor: Bradleya 5: 94. (1987) |
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