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netted nutrush, netted nutsedge

Muehlenberg's nutrush

Habit Plants; roots purplish, fibrous; rhizomes short (in perennial forms). Plants annual or sometimes perennial; roots fibrous; rhizomes present in perennial forms, short.
Culms

solitary or in tufts, erect or spreading, slender, (6–)15–50 cm, glabrous or hairy.

solitary or in tufts, erect or spreading, slender, 20–90(–125) cm, glabrous or hairy.

Leaves

sheaths often purple-tinged at base, sometimes somewhat winged, ribbed, glabrous or densely pubescent;

contra-ligules rotund-obtuse, short;

blades linear, flat, often ribbed, obtuse, shorter than or equaling culm, 1–3.5 mm wide, glabrous or occasionally margins and veins abaxially scabrous.

sheaths often purplish tinged at base, sometimes somewhat winged, ribbed, glabrous or densely pubescent;

contra-ligules rotund-obtuse, short;

blades linear, flat, often ribbed, obtuse, shorter than or equaling culm, 1–5(–8) mm wide, glabrous or sparsely hirsute, sometimes scabrous on margins and veins abaxially.

Inflorescences

axillary and terminal, paniculate;

panicles compact, 8–25 mm, lateral clusters sessile or on short erect peduncles to 20(–90) mm;

bracts subtending inflorescence leaflike, distalmost lateral bract 3/4 as long as to exceeding terminal internode.

axillary and terminal, paniculate;

panicles diffuse, 15–40(–80) mm, lateral clusters ± remote, the proximal pedunculate, peduncles often recurved or pendulous, filiform, compressed, (15–)20–100 mm;

bracts subtending inflorescence leaflike, a bracteal leaf subtending panicles, usually 1/4–3/4 length of terminal internode, margins scabrous.

Spikelets

bisexual or unisexual; staminate spikelets 2–3 mm, pistillate spikelets 3–4(–5) mm; staminate scales lanceolate; pistillate scales ovate-lanceolate, apex acuminate.

bisexual or unisexual; staminate spikelets 2–3 mm, pistillate spikelets 3–4(–5) mm; staminate scales lanceolate, pistillate scales ovate-lanceolate, acuminate.

Achenes

gray or brownish and/or with dark interangular lines and little pubescence, globose to ovoid, 1.5–2 mm, apex umbonate, surface reticulate to completely smooth, somewhat verrucose in some forms, reticulations pubescent with tufts of yellowish hairs;

hypogynium distinctly 3-lobed, lobes ovate-lanceolate, apex obtuse, sometimes emarginate.

gray or brownish and/or with dark interangular lines and little pubescence, globose to ovoid, 1–3(–3.5) mm, reticulate to completely smooth, somewhat verrucose in some forms, reticulations smooth or pubescent with tufts of yellowish hairs, apex umbonate;

hypogynium distinctly 3-lobed, lobes ovate-lanceolate, obtuse or subacute, sometimes cuspidate, appressed to base of achene.

Scleria reticularis

Scleria muehlenbergii

Phenology Fruiting summer–fall. Fruiting summer–fall.
Habitat Moist or wet sandy or sandy-peaty soil of pond and lake margins, wet savannas, and moist swales Moist to wet (rarely dry) pinelands, savannas, prairies, and bogs: 0–500 m
Elevation 0–500 m (0–1600 ft)
Distribution
from FNA
AL; CT; DE; FL; GA; IN; MA; MD; MI; MS; NC; NH; NJ; NY; PA; RI; SC; TX; VA; WI
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; LA; MD; MO; MS; NC; NJ; NY; OK; PA; SC; TN; TX; VA; Mexico; Central America; West Indies; n South America
[WildflowerSearch map]
[BONAP county map]
Discussion

Scleria reticularis is a smaller plant and less widely distributed than S. muehlenbergii. In addition to the difference noted in the key, S. reticularis differs from S. muehlenbergii in having the lobe of the hypogynium obtuse and emarginate at the apex rather than obtuse to acute, the terminal internode of the stem usually 3–8 cm rather than 6–30 cm, and scales of pistillate flowers mostly 3–4 mm rather than 4–5 mm.

Scleria reticularis is a relatively uniform species throughout its restricted range. It does not occur outside of the United States and is infrequent in the southern portion of its range. The key characters used to separate S. reticularis from S. muehlenbergii apply only to temperate collections.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Scleria muehlenbergii is uncommon inland from the Coastal Plain, occurring usually in wet places at lower elevations.

This complex in Scleria is a difficult one. Recent authors recognized this plant as only a variety of S. reticularis, using the name S. reticularis var. pubescens. Taking all characters into consideration, S. muehlenbergii appears to be geographically, ecologically, and morphologically distinct from S. reticularis in the flora. As recognized here, S. muehlenbergii is quite variable in stature and achene size and sculpturing. In the United States, occasional plants occur with glabrous achenes, and some collections have achenes lacking or almost lacking sculpturing. Those variants maintain other features of S. muehlenbergii. Some tropical collections of the complex appear to be identical to United States material of S. muehlenbergii; other specimens appear somewhat different and combine characteristics in alternative ways. Those plants do not appear to be simple intermediates between S. muehlenbergii and S. reticularis. Plants from the Caribbean appear to be essentially identical to plants from the United States. Specimens of this complex from Mexico and Central America are also similar in most features to S. muehlenbergii, but some western Mexican plants are quite different, as noted by R. McVaugh (1993b), and Mexican and Central American plants usually have longer inflorescence bracts and often less pubescent achenes. More study is clearly needed here.

R. McVaugh (1993b) suggested that the oldest name available at varietal rank is Scleria bracteata var. angusta Grisebach. The combination under S. reticularis appears not to have been made.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 251. FNA vol. 23, p. 250.
Parent taxa Cyperaceae > Scleria Cyperaceae > Scleria
Sibling taxa
S. baldwinii, S. ciliata, S. curtissii, S. distans, S. georgiana, S. lacustris, S. lithosperma, S. minor, S. muehlenbergii, S. oligantha, S. pauciflora, S. triglomerata, S. verticillata
S. baldwinii, S. ciliata, S. curtissii, S. distans, S. georgiana, S. lacustris, S. lithosperma, S. minor, S. oligantha, S. pauciflora, S. reticularis, S. triglomerata, S. verticillata
Synonyms S. reticularis var. pubescens
Name authority Michaux: Fl. Bor.-Amer. 2: 167. (1803) Steudel: Nomencl. Bot. ed. 2, 2: 543. (1841)
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