Scleria oligantha |
Scleria |
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littlehead nutrush |
nutrush |
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Habit | Plants perennial; rhizomes elongate, nodulose, to 5 mm thick. | Herbs usually perennial, sometimes annual, cespitose or not, rhizomatous or not. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Culms | usually in tufts, erect, slender, 30–60(–70) cm, sometimes appearing weak, glabrous or hairy. |
solitary or not, unbranched proximal to inflorescence, trigonous, (6–)10–100(–125) cm (high-climbing into trees), weak or wiry. |
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Leaves | proximal sheaths purple tinged, the distal rosy red-tinged, usually narrowly winged, ribbed, pubescent or glabrous; contra-ligules ovate; blades linear, ribbed, shorter than or equaling culms, 2–6 mm wide, glabrous or ciliate to scabrous. |
few to many per culm, cauline, 3-ranked; sheaths present; ligules usually well developed, sometimes obsolete; blades well developed (or rudimentary, sometimes absent on proximal leaves), shorter than, equaling, or overtopping culms, plane, keeled, or revolute, 0.5–9 mm wide, stiff, with prominent midvein, glabrous, scabrous, or pubescent. |
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Inflorescences | axillary and terminal, of 2–5 small clusters, 5–20 × 5–10 mm, each with 1–4 spikelets; the lateral on reddish filiform peduncles dilated toward apex, hairy or scabrous on angles; bracts subtending inflorescence leaflike, linear-lanceolate, 3–10 cm, glabrous or ciliate. |
terminal, often axillary, sometimes pseudolateral, panicles, spikes, or 1 or more capitate or fasciculate clusters; spikelets 1–100+; bracts 1–3, ascending or erect, bristle-shaped, awl-shaped, or leaflike, sometimes appearing to be continuation of culm, 10–150 mm, shorter than or exceeding spikelets, glabrous or ciliate. |
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Spikelets | bisexual and staminate, staminate part frequently reduced in bisexual spikelets, purple tinged, 3–8 mm; staminate scales lanceolate; pistillate scales ovate, acuminate. |
scales 3–10+, each subtending flower, distichous, proximal 2–4 scales empty, proximal fertile scale pistillate, distal fertile scales staminate or sometimes empty; occasionally some spikelets entirely pistillate or staminate. |
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Flowers | unisexual; perianth absent; stamens 1–3; anthers 2–4 mm, apex usually mucronate or awned; styles linear, 2–3-fid, base deciduous or persistent, hypogynium rudimentary or prominent, slender or enlarged. |
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Achenes | white or grayish or grayish brown with dark longitudinal bands, ovoid, shorter than scales, 3–4 mm, smooth, shining, apex umbonate; hypogynium golden brown, rather broad, base obtuse-trigonous, supporting 8–9 small, round or elongate, granulose-spiculose tubercles. |
trigonous or rounded, 1–4 mm, smooth, tuberculate, verrucose, or reticulate, occasionally pubescent. |
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Scleria oligantha |
Scleria |
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Phenology | Fruiting spring–summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Mixed mesic to xeric woods or glades, wet meadows | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0–500 m (0–1600 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AL; AR; DC; FL; GA; IL; IN; KY; LA; MO; MS; NC; OH; OK; SC; TN; TX; VA; Mexico; Central America (Guatemala, Honduras)
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Tropical and warm-temperate regions of both hemispheres |
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Discussion | Scleria oligantha sometimes extends from its typical wooded habitat to open areas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 200 (14 in the flora). Unusual structures provide important taxonomic markers in Scleria, necessitating some specialized terminology. The hypogynium is a peculiar, hardened disc at the base of the achene. In some species the hypogynium forms a prominent collar, while in others it is an inconspicuous ridge around the point of attachment of the achene. Before the achene abscises, the hypogynium is attached to a concave disc, the cupula, which remains with the inflorescence when the achene is shed. The contra-ligule (sometimes called the ligula) is a membranous flap on the rim of the leaf sheath on the opposite side from the blade. Contra-ligules are present in some other genera of Cyperaceae; their structure is seldom used taxonomically outside Scleria. Scleria vaginata Steudel, native to Central and South America, was collected once as an adventive in southern Florida, where it is an aggressive vine over two meters. The hypogynium is well developed but lacks tubercles, and the achene is smooth, shining, and gray or purplish. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 247. | FNA vol. 23, p. 242. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Cyperaceae > Scleria | Cyperaceae | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Michaux: Fl. Bor.-Amer. 2: 167. (1803) | P. J. Bergius: Kongl. Vetensk. Acad. Handl. 26: 142, plates 4, 5. (1765) | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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