Scleria |
Scleria triglomerata |
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nutrush |
whip nutrush, whip nutsedge |
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Habit | Herbs usually perennial, sometimes annual, cespitose or not, rhizomatous or not. | Plants perennial; rhizomes clustered, stout, nodulose, hard. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Culms | solitary or not, unbranched proximal to inflorescence, trigonous, (6–)10–100(–125) cm (high-climbing into trees), weak or wiry. |
usually in tufts, stout, 40–100 cm, glabrous, somewhat scabrous distally. |
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Leaves | few to many per culm, cauline, 3-ranked; sheaths present; ligules usually well developed, sometimes obsolete; blades well developed (or rudimentary, sometimes absent on proximal leaves), shorter than, equaling, or overtopping culms, plane, keeled, or revolute, 0.5–9 mm wide, stiff, with prominent midvein, glabrous, scabrous, or pubescent. |
sheaths purplish, not winged, pilose or glabrous; contra-ligules ovate, rarely trigonous; blades linear, ribbed, shorter than culms, 3–9 mm wide, rigid, margins and midrib scabrous, sometimes slightly pubescent. |
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Inflorescences | terminal, often axillary, sometimes pseudolateral, panicles, spikes, or 1 or more capitate or fasciculate clusters; spikelets 1–100+; bracts 1–3, ascending or erect, bristle-shaped, awl-shaped, or leaflike, sometimes appearing to be continuation of culm, 10–150 mm, shorter than or exceeding spikelets, glabrous or ciliate. |
terminal and axillary, fasciculate; fascicles 3, 15–40 × 5–15 mm, each with (1–) 3–10(–12) spikelets (smaller numbers in shade forms); lateral peduncles erect, often becoming filiform and pendulous in deep shade forms; bracts subtending inflorescence leaflike, lanceolate, 3–11 cm, long-acuminate-attenuate, ciliate or glabrous. |
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Spikelets | scales 3–10+, each subtending flower, distichous, proximal 2–4 scales empty, proximal fertile scale pistillate, distal fertile scales staminate or sometimes empty; occasionally some spikelets entirely pistillate or staminate. |
bisexual and staminate (sometimes reduced to few flowers in bisexual spikelets), brown, 3–9 mm; staminate scales ovate-mucronate to lanceolate-acuminate, pistillate scales ovate, midrib excurrent, often awnlike. |
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Flowers | unisexual; perianth absent; stamens 1–3; anthers 2–4 mm, apex usually mucronate or awned; styles linear, 2–3-fid, base deciduous or persistent, hypogynium rudimentary or prominent, slender or enlarged. |
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Achenes | trigonous or rounded, 1–4 mm, smooth, tuberculate, verrucose, or reticulate, occasionally pubescent. |
sometimes grayish brown, sometimes with dark longitudinal bands, ovoid to subglobose, 2–3 mm, smooth, shining, apex obtuse; hypogynium low, obscurely 3-angled, covered with whitish or brownish, siliceous, papillose-spiculose crust. |
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Scleria |
Scleria triglomerata |
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Phenology | Fruiting summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Dark woods to open, moist pinelands, meadows, swales, prairies, and savannas | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0–900 m (0–3000 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
Tropical and warm-temperate regions of both hemispheres |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON
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Discussion | Species ca. 200 (14 in the flora). Unusual structures provide important taxonomic markers in Scleria, necessitating some specialized terminology. The hypogynium is a peculiar, hardened disc at the base of the achene. In some species the hypogynium forms a prominent collar, while in others it is an inconspicuous ridge around the point of attachment of the achene. Before the achene abscises, the hypogynium is attached to a concave disc, the cupula, which remains with the inflorescence when the achene is shed. The contra-ligule (sometimes called the ligula) is a membranous flap on the rim of the leaf sheath on the opposite side from the blade. Contra-ligules are present in some other genera of Cyperaceae; their structure is seldom used taxonomically outside Scleria. Scleria vaginata Steudel, native to Central and South America, was collected once as an adventive in southern Florida, where it is an aggressive vine over two meters. The hypogynium is well developed but lacks tubercles, and the achene is smooth, shining, and gray or purplish. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Scleria triglomerata is by far the most common and widespread species of Scleria in the flora region. It exhibits considerable ecologic amplitude, growing in shady woods to open pinelands. Past authors have sometimes separated the species into several, recognizing S. nitida and S. flaccida. Plants that fit all the characteristics of the segregates seem distinctive; apparent intermediates occur. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 242. | FNA vol. 23. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | S. flaccida, S. nitida | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | P. J. Bergius: Kongl. Vetensk. Acad. Handl. 26: 142, plates 4, 5. (1765) | Michaux: Fl. Bor.-Amer. 2: 168. (1803) | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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