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dark-green bulrush, green bulrush, scirpe noirâtre, woolgrass bulrush

wood bulrush, woodland bulrush

Habit Plants cespitose; rhizomes short, tough, fibrous. Plants spreading; rhizomes reddish, long, with conspicuous nodes and internodes.
Culms

fertile ones upright or nearly so;

nodes without axillary bulblets.

fertile ones upright or nearly so;

nodes without axillary bulblets.

Leaves

6–11 per culm;

sheaths of proximal leaves light brown;

proximal sheaths and blades with septa many, ± conspicuous;

blades 20–54 cm × 7–17 mm.

5–8 per culm;

sheaths of proximal leaves red;

proximal sheaths and blades with septa many, conspicuous;

blades 30–68 cm × 9–23 mm.

Inflorescences

terminal, rarely also with 1 lateral inflorescence from distal leaf axil;

rays ascending or divergent (commonly both in the same inflorescence), proximal branches smooth, distal branches scabrellous to scabrous, rays often bearing axillary bulblets;

bases of involucral bracts green, margins usually speckled with red-brown, rarely solid black, not glutinous.

terminal;

rays divaricate or occasionally ascending, proximal branches scabrellous (rarely smooth), distal branches scabrous, rays without axillary bulblets;

bases of involucral bracts green or reddish, not glutinous.

Spikelets

in dense clusters of 4–110 (largest cluster with 17–25+), spikelets sessile, ovoid to narrowly ovoid, 2–5(–8) × 1–2.5 mm;

scales dark brown with pale midribs, elliptic or broadly elliptic (rarely almost circular), 1.2–2.1 mm, apex mucronate, mucro 0.1–0.3(–0.4) mm.

in dense clusters of (1–)3–13(–24), (largest cluster with 7 or more), spikelets sessile, ovoid or narrowly ovoid, 2–6 × 1–3 mm;

scales black with green midribs, ovate or broadly rounded-ovate to nearly triangular or broadly triangular, 1–2.2 mm, apex rounded to mucronate, mucro (if present) to 0.2 mm.

Flowers

perianth bristles persistent, (5–)6, rather stout to somewhat slender, straight or curved, shorter than to slightly exceeding achene, with retrorse, thin-walled, round-tipped barbs in distal 0.3–0.6, enclosed within scales;

styles 3-fid.

perianth bristles brittle-based, readily detached from achene, 6, stout, straight or curved, shorter than to 1.5 times as long as achene, with retrorse, thick-walled, sharp-pointed teeth densely arranged almost to base, at maturity projecting beyond scales;

styles 3-fid.

Achenes

light brown, elliptic or obovate in outline, plumply trigonous or plano-convex, (0.8–)1–1.3 × 0.4–0.6 mm.

pale brown, oblong-elliptic to broadly elliptic or obovate in outline, plano-convex or sometimes plumply trigonous, 1–1.6 × (0.6–)0.8–1 mm.

2n

= 50–60.

= 64.

Scirpus atrovirens

Scirpus expansus

Phenology Fruiting late spring–early summer (late Jun–Jul). Fruiting summer (Jul–Aug).
Habitat Marshes, moist meadows Marshes, wet meadows
Elevation 0–900 m (0–3000 ft) 0–1100 m (0–3600 ft)
Distribution
from FNA
AL; AZ; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SD; TN; TX; VA; VT; WI; WV; MB; NF; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; DE; GA; MA; MD; ME; MI; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WV; NS; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Scirpus atrovirens is variable and often confused with all of its close relatives: S. georgianus, S. flaccidifolius, S. hattorianus, and S. pallidus. The septa in the blades and sheaths of the proximal leaves tend to be more prominent in S. atrovirens than in any of the related species, so that the leaves and sheaths appear distinctly nodulose in dried specimens. The scales have relatively little black pigmentation, so that the heads appear brownish, not blackish as in S. pallidus, S. hattorianus, and S. flaccidifolius.

Scirpus atrovirens occasionally hybridizes with S. georgianus, S. hattorianus, S. pallidus, and S. polyphyllus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

North American reports of Scirpus sylvaticus Linnaeus, a widespread Eurasian species, are based on specimens of S. expansus and S. microcarpus.

Scirpus expansus occasionally hybridizes with S. microcarpus, particularly in New England, and the names S. sylvaticus var. bissellii Fernald, S. expansus forma bissellii (Fernald) Fernald, and S. rubrotinctus forma radiosus Fernald are based on specimens of S. expansus × microcarpus. Hybrids are easily recognized by their elongate spikelets in which most of the ovaries are empty and abortive. Despite the low fertility of these hybrids, hybrid populations are able to persist in some places, particularly south of the range of S. microcarpus in the Delaware Valley.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 15. FNA vol. 23, p. 17.
Parent taxa Cyperaceae > Scirpus Cyperaceae > Scirpus
Sibling taxa
S. ancistrochaetus, S. atrocinctus, S. congdonii, S. cyperinus, S. diffusus, S. divaricatus, S. expansus, S. flaccidifolius, S. georgianus, S. hattorianus, S. lineatus, S. longii, S. microcarpus, S. pallidus, S. pedicellatus, S. pendulus, S. polyphyllus
S. ancistrochaetus, S. atrocinctus, S. atrovirens, S. congdonii, S. cyperinus, S. diffusus, S. divaricatus, S. flaccidifolius, S. georgianus, S. hattorianus, S. lineatus, S. longii, S. microcarpus, S. pallidus, S. pedicellatus, S. pendulus, S. polyphyllus
Synonyms S. atrovirens var. pycnocephalus
Name authority Willdenow: Enum. Pl. 1: 79. (1809) Fernald: Rhodora 45: 293, plate 767, figs. 1–3. (1943)
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