Schoenoplectus tabernaemontani
Schoenoplectus mucronatus
great bulrush, scirpe des étangs, soft-stem bulrush, soft-stem club-bulrush, tule
bog bulrush, rice-field bulrush, rough-seed bulrush
3–10 mm diam.
cylindric, 0.5–3 m × 2–10 mm.
sharply trigonous, 0.4–1 m × 2–3 mm.
3–4, basal;
sheath fronts membranous-translucent, often pinnate-fibrillose;
blades 1–2, C-shaped to dorsiventrally flat in cross section, usually much shorter than sheath, distal blade 2–200 × 1–4 mm, margins often scabridulous.
1–2;
sheath fronts not pinnate-fibrillose;
ligules absent;
blades absent.
2–4 times branched, branches to 15(–25) cm;
proximal bract usually erect, thickly C-shaped to subterete, 1–8 cm, margins sometimes scabridulous.
capitate;
proximal bract divergent to reflexed or rarely erect, trigonous, adaxially channeled, 1–10 cm.
15–200, solitary or in clusters of 2–4(–7), commonly all solitary, 3–17 × 2.5–4 mm;
scales uniformly dark to pale orange-brown, sometimes straw-colored, sometimes prominently lineolate-spotted, midrib often pale or green, ovate, 2–3.5 × 1.5–2 mm, sparsely (rarely densely) reddish or straw-colored, scabrous on awn and distal parts of midrib and sometimes flanks, margins ciliate, hairs contorted;
flanks veinless, apex obtuse to rounded, notch 0.2–0.3 mm deep, awn straight or bent, 0.2–0.8 mm.
4–20, 7–12 × 4 mm;
scales orange-brown to straw-colored, central region often greenish, broadly obovate, 3–3.5 × 2–2.5 mm, margins ciliolate, apex obtuse to broadly acute, entire, mucronate.
perianth bristles 6, brown, ± equaling achene, densely retrorsely spinulose;
anthers 2 mm;
styles 2-fid, sometimes 3-fid near spikelet apex.
perianth members 6, brown, bristlelike, equaling achene, stout, retrorsely spinulose;
anthers 0.8 mm;
styles 3-fid (or some 2-fid in same spikelet).
dark gray-brown when ripe, plano-convex, obovoid, 1.5–2.8 × 1.2–1.7 mm;
beak 0.2–0.4 mm.
dark to blackish brown, thickly plano-convex to obtusely trigonous, broadly obovoid, 1.7–2.2(–2.5) × 1.2–1.7 mm;
beak 0.2 mm.
= 42.
Schoenoplectus tabernaemontani
Schoenoplectus mucronatus
Two yellow-striped forms of Schoenoplectus tabernaemontani are grown as ornamentals.
Schoenoplectus validus, described from the Caribbean, and S. tabernaemontani, described from Europe, are here treated as one variable, cosmopolitan species without infraspecific taxa, pending further studies (J. Browning et al. 1995b; S. G. Smith 1995). Most North American plants have spikelets with reddish papillae or prickles on the scales, whereas some plants of coastal and boreal North America closely resemble most plants of northwestern Europe and southern Africa in their densely reddish prickly-papillose scales and are similar to the type of Scirpus glaucus J. E. Smith.
Schoenoplectus tabernaemontani, S. acutus, S. heterochaetus, S. lacustris, and S. triqueter belong to the very difficult S. lacustris complex. The entire complex except S. triqueter was treated as the single species Scirpus lacustris (T. Koyama 1962b). Many Old World authors treat Schoenoplectus tabernaemontani as S. lacustris var. tabernaemontani or subsp. glaucus.
Much of the local infraspecific variation in the Schoenoplectus lacustris complex is probably because of hybridization. Some studies support the recognition of separate species in this group (J. Browning et al. 1995b). Hybrids in North America include S. acutus × S. tabernaemontani, widespread and common, especially in the east; S. acutus × S. heterochaetus = S. ×oblongus (T. Koyama) Soják, widespread but uncommon; S. heterochaetus × S. tabernaemontani = S. ×steinmetzii (Fernald) S. G. Smith, eastern and most uncommon; S. tabernaemontani × S. triqueter = S. ×kuekenthalianus (Junge) Kent, lower Columbia River in Oregon and probably Washington; and S. acutus var. occidentalis × S. californicus, local in California. Except for its trigonous culms, S. triqueter is very similar to the S. lacustris complex and freely hybridizes with S. tabernaemontani, both in North America and Europe.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Schoenoplectus mucronatus was collected before 1900 in New Brunswick and New Jersey; apparently the plants did not persist. It has also been reported from New York and Pennsylvania; I have not seen specimens. Elsewhere, it has become firmly established. It is an important ricefield weed in California (M. K. Bellue 1947), where it was first observed in 1942 and is called “ricefield bulrush.” It was first observed in the Midwest in 1971. Schoenoplectus mucronatus is cultivated for wildlife food near the Columbia River in Clark County, Washington, but apparently is not established in that area. Schoenoplectus mucronatus is very similar to S. triangulatus (Roxburgh) Soják of Asia, which differs in its larger spikelets, spikelet scales, and anthers.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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