Schoenoplectus tabernaemontani
Schoenoplectus californicus
great bulrush, scirpe des étangs, soft-stem bulrush, soft-stem club-bulrush, tule
California bulrush, giant bulrush, southern bulrush, tule
3–10 mm diam.
10–15 mm diam.
cylindric, 0.5–3 m × 2–10 mm.
bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth.
3–4, basal;
sheath fronts membranous-translucent, often pinnate-fibrillose;
blades 1–2, C-shaped to dorsiventrally flat in cross section, usually much shorter than sheath, distal blade 2–200 × 1–4 mm, margins often scabridulous.
3–4, basal, ca. 1/6 culm length;
sheath fronts coarsely pinnate-fibrillose;
blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm.
2–4 times branched, branches to 15(–25) cm;
proximal bract usually erect, thickly C-shaped to subterete, 1–8 cm, margins sometimes scabridulous.
3 or more times branched, branches to 21 cm;
proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous.
15–200, solitary or in clusters of 2–4(–7), commonly all solitary, 3–17 × 2.5–4 mm;
scales uniformly dark to pale orange-brown, sometimes straw-colored, sometimes prominently lineolate-spotted, midrib often pale or green, ovate, 2–3.5 × 1.5–2 mm, sparsely (rarely densely) reddish or straw-colored, scabrous on awn and distal parts of midrib and sometimes flanks, margins ciliate, hairs contorted;
flanks veinless, apex obtuse to rounded, notch 0.2–0.3 mm deep, awn straight or bent, 0.2–0.8 mm.
25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm;
scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous.
perianth bristles 6, brown, ± equaling achene, densely retrorsely spinulose;
anthers 2 mm;
styles 2-fid, sometimes 3-fid near spikelet apex.
perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs;
anthers 1.5 mm;
styles 2-fid.
dark gray-brown when ripe, plano-convex, obovoid, 1.5–2.8 × 1.2–1.7 mm;
beak 0.2–0.4 mm.
dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm;
beak 0.2 mm.
= 42.
= 68.
Schoenoplectus tabernaemontani
Schoenoplectus californicus
Two yellow-striped forms of Schoenoplectus tabernaemontani are grown as ornamentals.
Schoenoplectus validus, described from the Caribbean, and S. tabernaemontani, described from Europe, are here treated as one variable, cosmopolitan species without infraspecific taxa, pending further studies (J. Browning et al. 1995b; S. G. Smith 1995). Most North American plants have spikelets with reddish papillae or prickles on the scales, whereas some plants of coastal and boreal North America closely resemble most plants of northwestern Europe and southern Africa in their densely reddish prickly-papillose scales and are similar to the type of Scirpus glaucus J. E. Smith.
Schoenoplectus tabernaemontani, S. acutus, S. heterochaetus, S. lacustris, and S. triqueter belong to the very difficult S. lacustris complex. The entire complex except S. triqueter was treated as the single species Scirpus lacustris (T. Koyama 1962b). Many Old World authors treat Schoenoplectus tabernaemontani as S. lacustris var. tabernaemontani or subsp. glaucus.
Much of the local infraspecific variation in the Schoenoplectus lacustris complex is probably because of hybridization. Some studies support the recognition of separate species in this group (J. Browning et al. 1995b). Hybrids in North America include S. acutus × S. tabernaemontani, widespread and common, especially in the east; S. acutus × S. heterochaetus = S. ×oblongus (T. Koyama) Soják, widespread but uncommon; S. heterochaetus × S. tabernaemontani = S. ×steinmetzii (Fernald) S. G. Smith, eastern and most uncommon; S. tabernaemontani × S. triqueter = S. ×kuekenthalianus (Junge) Kent, lower Columbia River in Oregon and probably Washington; and S. acutus var. occidentalis × S. californicus, local in California. Except for its trigonous culms, S. triqueter is very similar to the S. lacustris complex and freely hybridizes with S. tabernaemontani, both in North America and Europe.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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