Schoenoplectus californicus |
Schoenoplectus torreyi |
|
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California bulrush, giant bulrush, southern bulrush, tule |
scirpe de Torrey, Torrey's bulrush, Torrey's club-bulrush, Torrey's three-square bulrush |
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Habit | Plants mat-forming; rhizomes 1–3 mm diam. | |
Rhizomes | 10–15 mm diam. |
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Culms | bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth. |
sharply trigonous, sides concave distally, 0.4–1.5 m × 1.5–5 mm. |
Leaves | 3–4, basal, ca. 1/6 culm length; sheath fronts coarsely pinnate-fibrillose; blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm. |
4–7, nearly equaling culm; sheath fronts delicately pinnate-fibrillose; blades 3–6, erect, the distal mostly longer than sheath, 1–2 mm, wide, proximally thickly V-shaped central region trigonous-channeled to laterally flattened, apically asymmetically laterally flattened-trigonous; apex often markedly eccentric, glabrous, smooth or angles papillose. |
Inflorescences | 3 or more times branched, branches to 21 cm; proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous. |
capitate or 1 spikelet; proximal bract erect, resembling leaf blade, 3–20 cm. |
Spikelets | 25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm; scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous. |
1–4, 7–18 × 3–5 mm; scales straw-colored to orange-brown or central region greenish, ovate-lanceolate, 4–5 × 2.5 mm, margins entire, smooth or awn sparsely spinulose, sometimes distally ciliolate at 40X, apex acute, mucronate, mucro 0.2–0.5 mm. |
Flowers | perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs; anthers 1.5 mm; styles 2-fid. |
perianth members 6, brown, bristlelike, equaling to slightly exceeding achene, spinulose; anthers 2–3 mm; styles 3-fid. |
Achenes | dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm; beak 0.2 mm. |
brown, compressed trigonous, ovoid to obovoid, 3.5–4.5 × 1.7–2 mm; beak 0.5–1 mm. |
2n | = 68. |
= 70. |
Schoenoplectus californicus |
Schoenoplectus torreyi |
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Phenology | Fruiting spring–summer. | Fruiting summer. |
Habitat | Brackish to fresh marshes, shores, often emergent in water | Emergent in fresh ponds and marshes, often with fluctuating water levels |
Elevation | 0–1400 m [0–4600 ft] | 10–200 m [30–700 ft] |
Distribution |
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
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CT; DE; IA; IL; IN; KY; MA; ME; MI; MN; MO; NH; NJ; NY; OH; PA; RI; VA; VT; WI; MB; NB; ON; QC
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Discussion | Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Schoenoplectus torreyi is very locally distributed, and probably has been extirpated from many historic localities. A specimen from an uncertain locality in Kentucky was collected in 1830. I have not seen specimens from North Dakota, South Dakota, or Nebraska. Scirpus torreyi was placed in synonymy under S. subterminalis Torrey var. cylindricus (Torrey) T. Koyama [misapplied, = Bolboschoenus novae-angliae by T. Koyama (1962b)]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 50. | FNA vol. 23, p. 54. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Elytrospermum californicum, Scirpus californicus | Scirpus torreyi |
Name authority | (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) | (Olney) Palla: Allg. Bot. Z. Syst. 17(Beil.): 3. (1912) |
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