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California bulrush, giant bulrush, southern bulrush, tule

scirpe de Torrey, Torrey's bulrush, Torrey's club-bulrush, Torrey's three-square bulrush

Habit Plants mat-forming; rhizomes 1–3 mm diam.
Rhizomes

10–15 mm diam.

Culms

bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth.

sharply trigonous, sides concave distally, 0.4–1.5 m × 1.5–5 mm.

Leaves

3–4, basal, ca. 1/6 culm length;

sheath fronts coarsely pinnate-fibrillose;

blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm.

4–7, nearly equaling culm;

sheath fronts delicately pinnate-fibrillose;

blades 3–6, erect, the distal mostly longer than sheath, 1–2 mm, wide, proximally thickly V-shaped central region trigonous-channeled to laterally flattened, apically asymmetically laterally flattened-trigonous;

apex often markedly eccentric, glabrous, smooth or angles papillose.

Inflorescences

3 or more times branched, branches to 21 cm;

proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous.

capitate or 1 spikelet;

proximal bract erect, resembling leaf blade, 3–20 cm.

Spikelets

25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm;

scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous.

1–4, 7–18 × 3–5 mm;

scales straw-colored to orange-brown or central region greenish, ovate-lanceolate, 4–5 × 2.5 mm, margins entire, smooth or awn sparsely spinulose, sometimes distally ciliolate at 40X, apex acute, mucronate, mucro 0.2–0.5 mm.

Flowers

perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs;

anthers 1.5 mm;

styles 2-fid.

perianth members 6, brown, bristlelike, equaling to slightly exceeding achene, spinulose;

anthers 2–3 mm;

styles 3-fid.

Achenes

dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm;

beak 0.2 mm.

brown, compressed trigonous, ovoid to obovoid, 3.5–4.5 × 1.7–2 mm;

beak 0.5–1 mm.

2n

= 68.

= 70.

Schoenoplectus californicus

Schoenoplectus torreyi

Phenology Fruiting spring–summer. Fruiting summer.
Habitat Brackish to fresh marshes, shores, often emergent in water Emergent in fresh ponds and marshes, often with fluctuating water levels
Elevation 0–1400 m [0–4600 ft] 10–200 m [30–700 ft]
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DE; IA; IL; IN; KY; MA; ME; MI; MN; MO; NH; NJ; NY; OH; PA; RI; VA; VT; WI; MB; NB; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Schoenoplectus torreyi is very locally distributed, and probably has been extirpated from many historic localities. A specimen from an uncertain locality in Kentucky was collected in 1830. I have not seen specimens from North Dakota, South Dakota, or Nebraska. Scirpus torreyi was placed in synonymy under S. subterminalis Torrey var. cylindricus (Torrey) T. Koyama [misapplied, = Bolboschoenus novae-angliae by T. Koyama (1962b)].

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 50. FNA vol. 23, p. 54.
Parent taxa Cyperaceae > Schoenoplectus > sect. Schoenoplectus Cyperaceae > Schoenoplectus > sect. Malacogeton
Sibling taxa
S. acutus, S. americanus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
S. acutus, S. americanus, S. californicus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. triqueter
Synonyms Elytrospermum californicum, Scirpus californicus Scirpus torreyi
Name authority (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) (Olney) Palla: Allg. Bot. Z. Syst. 17(Beil.): 3. (1912)
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