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California bulrush, giant bulrush, southern bulrush, tule

Rocky Mountain bulrush

Habit Plants annual or perennial; rhizomes 1 mm diam.
Rhizomes

10–15 mm diam.

Culms

bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth.

often arching to decumbent, cylindric, ridged when dry, 0.09–0.65 m × 0.5–1.5 mm.

Leaves

3–4, basal, ca. 1/6 culm length;

sheath fronts coarsely pinnate-fibrillose;

blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm.

3–4, 1(–2) cauline;

sheath fronts not pinnate-fibrillose;

blades 1–2, proximally thickly C-shaped in cross section, distally flat, rudimentary to longer than sheath, 1–200 × 0.2–1 mm, smooth or margins distally spinulose.

Inflorescences

3 or more times branched, branches to 21 cm;

proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous.

capitate or of 1 spikelet or with 1 or 2 branches to 15 mm;

proximal bract erect, like leaf blades, 5–15 cm.

Spikelet(s)

25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm;

scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous.

achenes blackish brown, nearly equilaterally sharply trigonous, ovoid to obovoid, 1.3–1.8 × 1–1.4 mm, with 10–20 mostly sharp ridges;

beak 0.1 mm.

Flowers

perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs;

anthers 1.5 mm;

styles 2-fid.

perianth absent;

anthers 0.3–0.5 mm;

styles 3-fid.

Achenes

dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm;

beak 0.2 mm.

Amphicarpic

achenes trigonous, 2.2–3 mm including stout, tapered beak 0.3 mm, with many obscure to evident horizontal ridges.

2n

= 68.

= 50.

Schoenoplectus californicus

Schoenoplectus saximontanus

Phenology Fruiting spring–summer. Fruiting late summer–fall, spring–fall (California, Texas).
Habitat Brackish to fresh marshes, shores, often emergent in water Damp soils to emergent, freshwater ponds, ditches, often drying, disturbed, and sandy areas
Elevation 0–1400 m [0–4600 ft] 0–2200 m [0–7200 ft]
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; IL; KS; MO; NE; OH; OK; SD; TX; UT; WY; BC; Mexico (San Luis Potosí, Tamaulipas)
[WildflowerSearch map]
Discussion

Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The distribution of Schoenoplectus saximontanus is very scattered (local).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 50. FNA vol. 23, p. 59.
Parent taxa Cyperaceae > Schoenoplectus > sect. Schoenoplectus Cyperaceae > Schoenoplectus > sect. Supini
Sibling taxa
S. acutus, S. americanus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
S. acutus, S. americanus, S. californicus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
Synonyms Elytrospermum californicum, Scirpus californicus Scirpus saximontanus, Scirpus bergsonii, Scirpus supinus var. saximontanus
Name authority (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) (Fernald) J. Raynal: Adansonia, n. s. 16: 141. (1976)
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