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California bulrush, giant bulrush, southern bulrush, tule

Hall's annual-bulrush, Hall's bulrush

Habit Plants annual; rhizomes 1 mm diam.
Rhizomes

10–15 mm diam.

Culms

bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth.

often arching to decumbent, cylindric, ridged when dry, 0.05–0.8 m × 0.5–1 mm.

Leaves

3–4, basal, ca. 1/6 culm length;

sheath fronts coarsely pinnate-fibrillose;

blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm.

3–4, 1 cauline;

sheaths straw-colored to brown and disintegrating to fibers, front translucent and splitting;

blades 1–2, proximally thickly C-shaped in cross section, distally flat, from much shorter to longer than sheath, 1–200+ × 0.2–1 mm, smooth or margins distally spinulose.

Inflorescences

3 or more times branched, branches to 21 cm;

proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous.

capitate or 1 spikelet or rarely with 1 or 2 branches to 12 mm;

proximal bract erect, resembling leaf blades, 5–150 mm.

Spikelet(s)

25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm;

scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous.

achenes blackish brown, plano-convex or usually the adaxial surface longitudinally concave, ovoid to obovoid, 1.3–1.7 × 1.2–1.3 mm, with 15–18 mostly sharp ridges;

beak 0.1 mm.

Flowers

perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs;

anthers 1.5 mm;

styles 2-fid.

perianth absent;

anthers 0.4 mm;

styles 2-fid.

Achenes

dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm;

beak 0.2 mm.

Amphicarpic

achenes thickly trigonous with convex abaxial face, 1.7–2.5 × 1.1–1.6 mm, rugose with rounded ridges;

beak 0.3 mm.

2n

= 68.

= 22.

Schoenoplectus californicus

Schoenoplectus hallii

Phenology Fruiting spring–summer. Fruiting summer–fall.
Habitat Brackish to fresh marshes, shores, often emergent in water Terrestrial to emergent, freshwater shores, temporary ponds, wet places in cultivated fields, pastures, ditches, sinkholes, prairie
Elevation 0–1400 m [0–4600 ft] 70–700 m [230–2300 ft]
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
[WildflowerSearch map]
[BONAP county map]
from FNA
GA; IL; IN; KS; KY; MA; MI; MO; NE; OK; WI
Discussion

Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Many reports of Schoenoplectus hallii are based on misidentified specimens of S. saximontanus or S. erectus. The Iowa specimen was probably collected in 1890. The species is probably extirpated from Massachusetts. I have identified a specimen from the Georgia coastal plain, where S. hallii and S. erectus are sympatric, as intermediate between the two.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 50. FNA vol. 23, p. 59.
Parent taxa Cyperaceae > Schoenoplectus > sect. Schoenoplectus Cyperaceae > Schoenoplectus > sect. Supini
Sibling taxa
S. acutus, S. americanus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
S. acutus, S. americanus, S. californicus, S. deltarum, S. erectus, S. etuberculatus, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
Synonyms Elytrospermum californicum, Scirpus californicus Scirpus hallii, Scirpus supinus var. hallii
Name authority (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) (A. Gray) S. G. Smith: Novon 5: 101. (1995)
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