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California bulrush, giant bulrush, southern bulrush, tule

common tule, hard-stem bulrush, hard-stem club-bulrush, schoenoplectus, viscous bulrush

Rhizomes

10–15 mm diam.

5–15 mm diam.

Culms

bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth.

cylindric, 1–4 m × 2–10 mm.

Leaves

3–4, basal, ca. 1/6 culm length;

sheath fronts coarsely pinnate-fibrillose;

blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm.

3–4, all basal;

sheaths often dark reddish proximally, front membranous-translucent and splitting, orifice adaxially deeply V-shaped;

blades 1–2, C-shaped to dorsiventrally flat in cross section, usually much shorter than sheath, distal blade 8–120 × 3–7 mm, margins often scabridulous.

Inflorescences

3 or more times branched, branches to 21 cm;

proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous.

2(–3) times branched, open or compact, branches 6(–18) cm;

proximal bract usually erect, thickly C-shaped in cross section, 1–9 cm, margins sometimes scabridulous.

Spikelets

25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm;

scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous.

3–190, solitary or in clusters of 2–8, never all solitary;

scales reddish to orange-brown to straw-colored, often variable on same scale, usually wholly or partly straw-colored and prominently lineolate-spotted at 10X, scale or midrib often green when young, ovate, 3–4 × 2–3 mm, sparsely to often densely reddish or straw-colored spinulose-papillose distally or on most of surface, margins ciliate, hairs long, contorted, flanks veinless, apex acute to obtuse, notch 0.3–0.5 mm deep, awn on at least some scales in spikelet usually strongly contorted, 0.5–2 mm (often broken off).

Flowers

perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs;

anthers 1.5 mm;

styles 2-fid.

perianth members (4–)6(–8), brown, bristlelike, equaling achene body or sometimes much shorter, rarely rudimentary, spinulose;

anthers 2 mm;

styles 2(–3)-fid.

Achenes

dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm;

beak 0.2 mm.

dark gray-brown, plano-convex or rarely weakly trigonous, obovoid, (1.5–)2–3  1.2–1.7 mm;

beak 0.2–0.4 mm.

2n

= 68.

Schoenoplectus californicus

Schoenoplectus acutus

Phenology Fruiting spring–summer.
Habitat Brackish to fresh marshes, shores, often emergent in water
Elevation 0–1400 m [0–4600 ft]
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WY; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK; YT; Mexico; possibly Eurasia; e Pacific Islands
[WildflowerSearch map]
[BONAP county map]
Discussion

Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Plants of Schoenoplectus acutus var. occidentalis that have most styles 3-fid, most achenes trigonous, and culms very soft with large air cavities occur at lower elevations in California (mostly southern) and Baja California. In California they often grow on stream bars. They may deserve varietal status. The varieties intergrade throughout most of the west with var. occidentalis.

Schoenoplectus acutus hybridizes with Schoenoplectus tabernaemontani, S. heterochaetus, and S. californicus (see also S. tabernaemontani).

Although previously assumed to be restricted to North America, some specimens from Eurasia are probably Schoenoplectus acutus, and one very young specimen from Clipperton Island in the eastern Pacific is probably S. acutus var. occidentalis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles 2-fid; achenes plano-convex; culms mostly very firm, air cavities in distal 1/4 mostly ca. 0.5 mm wide.
var. acutus
1. Some styles 3-fid; some achenes compressed, obtusely trigonous; culms firm to soft, larger air cavities in distal 1/4 1–2.5 mm wide.
var. occidentalis
Source FNA vol. 23, p. 50. FNA vol. 23, p. 48.
Parent taxa Cyperaceae > Schoenoplectus > sect. Schoenoplectus Cyperaceae > Schoenoplectus > sect. Schoenoplectus
Sibling taxa
S. acutus, S. americanus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
S. americanus, S. californicus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
Subordinate taxa
S. acutus var. acutus, S. acutus var. occidentalis
Synonyms Elytrospermum californicum, Scirpus californicus Scirpus acutus
Name authority (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) (Muhlenberg ex Bigelow) Á. Löve & D. Löve: Bull. Torrey Bot. Club 81: 33. (1954)
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