Schoenoplectus californicus |
Schoenoplectus subterminalis |
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California bulrush, giant bulrush, southern bulrush, tule |
scirpe subterminal, swaying bulrush, swaying clubrush, water annual-bulrush, water bulrush, water club-rush |
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Habit | Plants mat-forming; rhizomes to 1 mm diam; terminal tubers sometimes present, to 3 mm thick, fleshy. | |
Rhizomes | 10–15 mm diam. |
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Culms | bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth. |
subcylindric, 0.2–1.5 m × 0.5–1 mm. |
Leaves | 3–4, basal, ca. 1/6 culm length; sheath fronts coarsely pinnate-fibrillose; blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm. |
3–20+, smooth; sheath fronts sometimes delicately pinnate-fibrillose; blades 3–20, when erect C-shaped in cross section proximally, elliptic most of length, when submerged usually flaccid and filiform to narrowly ribbonlike, probably longer than sheath, 0.2–1 mm wide. |
Inflorescences | 3 or more times branched, branches to 21 cm; proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous. |
a solitary, erect spikelet; proximal bract erect, like stiff leaf blade, 7–60 mm. |
Spikelets | 25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm; scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous. |
5–15 × 3–7 mm; scales pale brown, or central region green when young, ovate-lanceolate, 4–6 × 1.5 mm, smooth, margins sometimes ciliolate apically at 40X, midrib obscure, apex acute, entire, minutely mucronate. |
Flowers | perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs; anthers 1.5 mm; styles 2-fid. |
perianth members 6, brown, bristlelike, from 1/2 of to equaling achene beak, spinulose; anthers 2–3.5 mm; styles 3-fid. |
Achenes | dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm; beak 0.2 mm. |
brown, strongly trigonous, obovoid to obpyriform, (2.5–)3–3.5 × 1.5–1.7 mm; beak 0.3–0.8 mm. |
2n | = 68. |
= 72. |
Schoenoplectus californicus |
Schoenoplectus subterminalis |
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Phenology | Fruiting spring–summer. | Fruiting summer. |
Habitat | Brackish to fresh marshes, shores, often emergent in water | Submerged to emergent in water to 1 m or sometimes terrestrial, fresh lakes, streams, bogs |
Elevation | 0–1400 m (0–4600 ft) | 10–2200 m (0–7200 ft) |
Distribution |
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
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AK; AL; CA; CT; DE; FL; GA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; NH; NJ; NY; OH; OR; PA; RI; SC; UT; VA; VT; WA; WI; BC; NB; NL; NS; ON; PE; QC; SK
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Discussion | Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Schoenoplectus subterminalis often forms lawnlike, underwater mats that are entirely vegetative or have only the inflorescences emergent. This species is probably extirpated from Illinois. Schoenoplectus subterminalis var. terrestris Paine [= S. subterminalis forma terrestris (Paine) Fernald] probably does not deserve taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 50. | FNA vol. 23, p. 54. |
Parent taxa | Cyperaceae > Schoenoplectus > sect. Schoenoplectus | Cyperaceae > Schoenoplectus > sect. Malacogeton |
Sibling taxa | ||
Synonyms | Elytrospermum californicum, Scirpus californicus | Scirpus subterminalis |
Name authority | (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) | (Torrey) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) |
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