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California bulrush, giant bulrush, southern bulrush, tule

scirpe subterminal, swaying bulrush, swaying clubrush, water annual-bulrush, water bulrush, water club-rush

Habit Plants mat-forming; rhizomes to 1 mm diam; terminal tubers sometimes present, to 3 mm thick, fleshy.
Rhizomes

10–15 mm diam.

Culms

bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth.

subcylindric, 0.2–1.5 m × 0.5–1 mm.

Leaves

3–4, basal, ca. 1/6 culm length;

sheath fronts coarsely pinnate-fibrillose;

blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm.

3–20+, smooth;

sheath fronts sometimes delicately pinnate-fibrillose;

blades 3–20, when erect C-shaped in cross section proximally, elliptic most of length, when submerged usually flaccid and filiform to narrowly ribbonlike, probably longer than sheath, 0.2–1 mm wide.

Inflorescences

3 or more times branched, branches to 21 cm;

proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous.

a solitary, erect spikelet;

proximal bract erect, like stiff leaf blade, 7–60 mm.

Spikelets

25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm;

scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous.

5–15 × 3–7 mm;

scales pale brown, or central region green when young, ovate-lanceolate, 4–6 × 1.5 mm, smooth, margins sometimes ciliolate apically at 40X, midrib obscure, apex acute, entire, minutely mucronate.

Flowers

perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs;

anthers 1.5 mm;

styles 2-fid.

perianth members 6, brown, bristlelike, from 1/2 of to equaling achene beak, spinulose;

anthers 2–3.5 mm;

styles 3-fid.

Achenes

dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm;

beak 0.2 mm.

brown, strongly trigonous, obovoid to obpyriform, (2.5–)3–3.5 × 1.5–1.7 mm;

beak 0.3–0.8 mm.

2n

= 68.

= 72.

Schoenoplectus californicus

Schoenoplectus subterminalis

Phenology Fruiting spring–summer. Fruiting summer.
Habitat Brackish to fresh marshes, shores, often emergent in water Submerged to emergent in water to 1 m or sometimes terrestrial, fresh lakes, streams, bogs
Elevation 0–1400 m (0–4600 ft) 10–2200 m (0–7200 ft)
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; CA; CT; DE; FL; GA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; NH; NJ; NY; OH; OR; PA; RI; SC; UT; VA; VT; WA; WI; BC; NB; NL; NS; ON; PE; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Schoenoplectus subterminalis often forms lawnlike, underwater mats that are entirely vegetative or have only the inflorescences emergent. This species is probably extirpated from Illinois. Schoenoplectus subterminalis var. terrestris Paine [= S. subterminalis forma terrestris (Paine) Fernald] probably does not deserve taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 50. FNA vol. 23, p. 54.
Parent taxa Cyperaceae > Schoenoplectus > sect. Schoenoplectus Cyperaceae > Schoenoplectus > sect. Malacogeton
Sibling taxa
S. acutus, S. americanus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
S. acutus, S. americanus, S. californicus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. tabernaemontani, S. torreyi, S. triqueter
Synonyms Elytrospermum californicum, Scirpus californicus Scirpus subterminalis
Name authority (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) (Torrey) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972)
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