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California bulrush, giant bulrush, southern bulrush, tule

pale great bulrush, scirpe à sois inégales, slender bulrush, slender club-bulrush

Rhizomes

10–15 mm diam.

5–8 mm diam.;

scales longer than internodes, disintegrating to fibers.

Culms

bluntly trigonous with nearly flat sides throughout to cylindric proximally, obscurely trigonous distally, 1–4 m × 4–10 mm, smooth.

cylindric, 1.5–2.5 m × 4–8 mm.

Leaves

3–4, basal, ca. 1/6 culm length;

sheath fronts coarsely pinnate-fibrillose;

blades 0–1, dorsiventrally flat, not more than 1/2 sheath length, distal blade to 20 × 2 mm.

basal;

sheaths often separating from culm, resembling blade, sometimes reddish proximally, front membranous-translucent, splitting, and delicately pinnate-fibrillose, orifice adaxially deeply V-shaped;

ligules entire, 1 mm;

blades 1–2, thinly C-shaped to thin and dorsiventrally flat in cross section, much shorter to longer than sheath, distal blade to 220 × 5 mm.

Inflorescences

3 or more times branched, branches to 21 cm;

proximal bract usually erect, obtusely trigonous to subcylindric-channeled, 1–8 cm, margins often scabridulous.

2x branched, open, plano- to concavo-convex, to 9 cm;

proximal bract usually erect, thickly C-shaped to subterete in cross section, 1–15 cm, margins often scabridulous.

Spikelets

25–150+, solitary and in clusters of 2 or more, 5–11 × 3 mm;

scales orange-brown, brightly lineolate-spotted, broadly oval, 2.5 × 2 mm, margins ciliolate, flanks veinless, apex rounded, notch 0.1–0.2 mm deep, awn not contorted, usually bent outward, 0.3 mm, sparsely scabrous.

5–30, solitary or rarely 2 together, 5–15 × 3–4 mm;

scales light orange-brown to nearly colorless, usually obscurely lineolate-spotted, midribs often greenish, ovate-oblong, 3–4 × 1.5 mm, sparsely reddish- or straw-colored-scabrous distally on awn, sometimes midrib, and rarely adjacent parts of sides, margins ciliate, hairs contorted, flanks veinless, apex obtuse, notch 0.5 mm deep, awn straight to slightly contorted, 0.3–1 mm.

Flowers

perianth members 2–4, dark red-brown, thickly strap-shaped or trigonous, equaling achenes, fringed with brownish, soft, blunt hairs;

anthers 1.5 mm;

styles 2-fid.

perianth members 4(–5), brown, bristlelike, equaling achene body or adaxial and abaxial much shorter, sparsely retrorsely spinulose;

anthers 2 mm;

styles 3-fid.

Achenes

dark gray-brown when ripe, unequally biconvex or plano-convex, ovoid to obovoid, 1.8–2.2 × 1.3 mm;

beak 0.2 mm.

dark gray-brown, compressed-trigonous, obovoid, 2.2–3.2 × 1.4–1.9 mm;

beak 0.3–1 mm.

2n

= 68.

= 38.

Schoenoplectus californicus

Schoenoplectus heterochaetus

Phenology Fruiting spring–summer. Fruiting summer.
Habitat Brackish to fresh marshes, shores, often emergent in water Fresh, often calcareous marshes and lakes, often emergent in water to 1.5 m
Elevation 0–1400 m (0–4600 ft) 10–500(–2000) m (0–1600(–6600) ft)
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; KS; LA; MS; NC; NM; NV; OK; SC; TN; TX; Mexico; Central America; West Indies; South America (s to Argentina, Chile); Pacific Islands (Cook Islands, Easter Island, Hawaii) [Introduced, New Zealand]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; IA; ID; IL; KS; MA; MI; MN; MO; MT; ND; NE; NY; OK; OR; PA; SD; VT; WI; WY; AB; MB; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Schoenoplectus californicus forms sterile hybrids with S. acutus in California (see 1. S. tabernaemontani) (S. G. Smith 1995). It reportedly occurs in Pope County, Illinois, although I have not seen a specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Schoenoplectus heterochaetus hybridizes with S. acutus and S. tabernaemontani (see comment under 1. S. tabernaemontani).

The Michigan record is dubious, the Oregon record is from 1895, and reports from Washington are apparently based on misidentified specimens. Some specimens from scattered localities in Eurasia may be Schoenoplectus heterochaetus.

Schoenoplectus lacustris (Linnaeus) Palla, of Eurasia, was collected near Camden, New Jersey, in 1889 and 1891; it now appears to be extirpated. It is also in cultivation in North America and should be sought. It is similar to S. heterochaetus, from which it differs in its often clustered spikelets, six long perianth bristles, and ability to form submerged ribbon-leaved forms, especially in flowing water in streams. 2n = 42.

Schoenoplectus lacustris hybridizes in Europe with S. tabernaemontani and S. triqueter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 50. FNA vol. 23, p. 49.
Parent taxa Cyperaceae > Schoenoplectus > sect. Schoenoplectus Cyperaceae > Schoenoplectus > sect. Schoenoplectus
Sibling taxa
S. acutus, S. americanus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. heterochaetus, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
S. acutus, S. americanus, S. californicus, S. deltarum, S. erectus, S. etuberculatus, S. hallii, S. mucronatus, S. pungens, S. purshianus, S. saximontanus, S. smithii, S. subterminalis, S. tabernaemontani, S. torreyi, S. triqueter
Synonyms Elytrospermum californicum, Scirpus californicus Scirpus heterochaetus, Scirpus lacustris var. tenuiculmis
Name authority (C. A. Meyer) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972) (Chase) Soják: Cas. Nár. Mus., Odd. Prír. 140: 127. (1972)
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