Schoenoplectus acutus |
Schoenoplectus mucronatus |
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common tule, hard-stem bulrush, hard-stem club-bulrush, schoenoplectus, viscous bulrush |
bog bulrush, rice-field bulrush, rough-seed bulrush |
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Habit | Plants perennial (or annual?); rhizomes hard. | |||||
Rhizomes | 5–15 mm diam. |
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Culms | cylindric, 1–4 m × 2–10 mm. |
sharply trigonous, 0.4–1 m × 2–3 mm. |
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Leaves | 3–4, all basal; sheaths often dark reddish proximally, front membranous-translucent and splitting, orifice adaxially deeply V-shaped; blades 1–2, C-shaped to dorsiventrally flat in cross section, usually much shorter than sheath, distal blade 8–120 × 3–7 mm, margins often scabridulous. |
1–2; sheath fronts not pinnate-fibrillose; ligules absent; blades absent. |
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Inflorescences | 2(–3) times branched, open or compact, branches 6(–18) cm; proximal bract usually erect, thickly C-shaped in cross section, 1–9 cm, margins sometimes scabridulous. |
capitate; proximal bract divergent to reflexed or rarely erect, trigonous, adaxially channeled, 1–10 cm. |
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Spikelets | 3–190, solitary or in clusters of 2–8, never all solitary; scales reddish to orange-brown to straw-colored, often variable on same scale, usually wholly or partly straw-colored and prominently lineolate-spotted at 10X, scale or midrib often green when young, ovate, 3–4 × 2–3 mm, sparsely to often densely reddish or straw-colored spinulose-papillose distally or on most of surface, margins ciliate, hairs long, contorted, flanks veinless, apex acute to obtuse, notch 0.3–0.5 mm deep, awn on at least some scales in spikelet usually strongly contorted, 0.5–2 mm (often broken off). |
4–20, 7–12 × 4 mm; scales orange-brown to straw-colored, central region often greenish, broadly obovate, 3–3.5 × 2–2.5 mm, margins ciliolate, apex obtuse to broadly acute, entire, mucronate. |
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Flowers | perianth members (4–)6(–8), brown, bristlelike, equaling achene body or sometimes much shorter, rarely rudimentary, spinulose; anthers 2 mm; styles 2(–3)-fid. |
perianth members 6, brown, bristlelike, equaling achene, stout, retrorsely spinulose; anthers 0.8 mm; styles 3-fid (or some 2-fid in same spikelet). |
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Achenes | dark gray-brown, plano-convex or rarely weakly trigonous, obovoid, (1.5–)2–3 1.2–1.7 mm; beak 0.2–0.4 mm. |
dark to blackish brown, thickly plano-convex to obtusely trigonous, broadly obovoid, 1.7–2.2(–2.5) × 1.2–1.7 mm; beak 0.2 mm. |
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Schoenoplectus acutus |
Schoenoplectus mucronatus |
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Phenology | Fruiting summer–fall. | |||||
Habitat | Wet soil to emergent in fresh water, ponds, ditches, rice fields | |||||
Elevation | 20–300 m (100–1000 ft) | |||||
Distribution |
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WY; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK; YT; Mexico; possibly Eurasia; e Pacific Islands
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CA; IA; IL; KY; MO; TN; Eurasia; Africa; Australia [Introduced in North America] |
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Discussion | Varieties 2 (2 in the flora). Plants of Schoenoplectus acutus var. occidentalis that have most styles 3-fid, most achenes trigonous, and culms very soft with large air cavities occur at lower elevations in California (mostly southern) and Baja California. In California they often grow on stream bars. They may deserve varietal status. The varieties intergrade throughout most of the west with var. occidentalis. Schoenoplectus acutus hybridizes with Schoenoplectus tabernaemontani, S. heterochaetus, and S. californicus (see also S. tabernaemontani). Although previously assumed to be restricted to North America, some specimens from Eurasia are probably Schoenoplectus acutus, and one very young specimen from Clipperton Island in the eastern Pacific is probably S. acutus var. occidentalis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Schoenoplectus mucronatus was collected before 1900 in New Brunswick and New Jersey; apparently the plants did not persist. It has also been reported from New York and Pennsylvania; I have not seen specimens. Elsewhere, it has become firmly established. It is an important ricefield weed in California (M. K. Bellue 1947), where it was first observed in 1942 and is called “ricefield bulrush.” It was first observed in the Midwest in 1971. Schoenoplectus mucronatus is cultivated for wildlife food near the Columbia River in Clark County, Washington, but apparently is not established in that area. Schoenoplectus mucronatus is very similar to S. triangulatus (Roxburgh) Soják of Asia, which differs in its larger spikelets, spikelet scales, and anthers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 48. | FNA vol. 23, p. 58. | ||||
Parent taxa | Cyperaceae > Schoenoplectus > sect. Schoenoplectus | Cyperaceae > Schoenoplectus > sect. Actaeogeton | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Scirpus acutus | Scirpus mucronatus | ||||
Name authority | (Muhlenberg ex Bigelow) Á. Löve & D. Löve: Bull. Torrey Bot. Club 81: 33. (1954) | (Linnaeus) Palla: Verh. K. K. Zool.-Bot. Ges. Wien 38(Sitzungsber.): 49. (1888) | ||||
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