Schistidium apocarpum |
Schistidium occidentale |
|
---|---|---|
schistidium moss |
western schistidium moss |
|
Habit | Plants in open tufts or mats, olivaceous to brownish (black), sometimes with yellowish tones. | Plants in open to occasionally compact tufts or mats, olivaceous, often (red-)brown or nearly black. |
Stems | 1.2–12 cm, central strand weak or absent. |
1–6 cm, sometimes denuded of leaves at base, central strand absent. |
Leaves | erect or curved, rarely (falcate-) secund when dry, ovate-lanceolate, sharply keeled distally, (1.3–)1.7–2.5(–3.2) mm, 1-stratose or rarely 2-stratose in striae distally; margins usually recurved throughout or to just before the apex, usually denticulate distally, 1 or 2-stratose; apices acute or sub-obtuse; costa percurrent or excurrent as a smooth or weakly denticulate, occasionally decurrent awn, abaxial surface often papillose; basal marginal cells usually quadrate; distal laminal cells mostly short-rectangular, 8–10 µm wide, smooth, sinuose. |
usually curved to falcate, sometimes falcate-secund or erect when dry, linear-lanceolate to ovate-lanceolate, concave proximally, weakly keeled or concave distally, 1.4–4(–6) mm, 1-stratose with a few 2-stratose striae or patches distally; margins plane or somewhat erect, rarely weakly recurved, smooth, 2-stratose; apices rounded or acute, usually ending in a fleshy, multistratose apiculus; costa percurrent, rarely short-excurrent, smooth; basal marginal cells quadrate or short-rectangular; distal cells quadrate or rounded, often angular, 6–11 µm wide, smooth, sometimes weakly bulging-mammillose, weakly sinuose. |
Sexual condition | autoicous. |
autoicous. |
Capsule | dark red or brown, short-cylindric, 0.7–1.3 mm; exothecial cells usually quadrate, sometimes mixed with short-elongate or oblate cells, thin-walled, usually trigonous; stomata present; peristome patent or erect, often twisted, 350–600(–700) µm, red, densely papillose, usually perforated. |
reddish brown, ovoid, cupulate, or short-cylindric, 1.3–2 mm; exothecial cells mostly short-elongate or isodiametric, usually irregularly angular, usually thin-walled, sometimes trigonous; rim darker colored than capsule wall, often red; stomata absent; peristome patent, 350–550 µm, red or orange-red, finely papillose, entire or perforated. |
Spores | 11–14(–19) µm, granulose or smooth. |
9–12 µm, smooth. |
Schistidium apocarpum |
Schistidium occidentale |
|
Phenology | Capsules mature late spring to early summer. | Capsules mature late spring to early summer. |
Habitat | Rocks in somewhat shaded habitats | Wet or dry rocks, often along intermittent watercourses |
Elevation | low to moderate elevations (0-1500 m) [low to moderate elevations (0-4900 ft)] | high elevations (2000-3500 m) [high elevations (6600-11500 ft)] |
Distribution |
AK; MI; NY; VT; WA; WI; AB; BC; NB; NF; NS; ON; QC; Greenland; Eurasia
|
CA; CO; ID; MT; NV; OR; UT; WA; WY |
Discussion | Although H. H. Blom (1996) considered Schistidium apocarpum to be restricted in the flora area to the eastern portions of North America, it is more widespread and scattered across the continent. The denticulate leaf margins, papillose abaxial costal surface, short-cylindrical capsules, and the thin-walled, often evenly quadrate exothecial cells are distinguishing characters. The long peristome teeth are also useful in identifying S. apocarpum. In good condition the peristome can only be confused with that of S. trichodon, which differs in the darker, often black color of its plants, the peristome teeth often forming a dome, and the cylindrical capsules that retain the columella. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
See comments under 6. Schistidium cinclidodonteum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 211. | FNA vol. 27, p. 221. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Grimmia apocarpa, S. lancifolium, S. umbrosum | Grimmia occidentalis |
Name authority | (Hedwig) Bruch & Schimper: Bryol. Europ. 3: 99. (1845) | (Lawton) Churchill: Advances Cladist. 1: 143. (1981) |
Web links |