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schistidium moss

Habit Plants in open tufts or mats, olivaceous to brownish (black), sometimes with yellowish tones. Plants in cushions, olivaceous, light brown, or green.
Stems

1.2–12 cm, central strand weak or absent.

0.5–2 cm, central strand indistinct.

Leaves

erect or curved, rarely (falcate-) secund when dry, ovate-lanceolate, sharply keeled distally, (1.3–)1.7–2.5(–3.2) mm, 1-stratose or rarely 2-stratose in striae distally;

margins usually recurved throughout or to just before the apex, usually denticulate distally, 1 or 2-stratose;

apices acute or sub-obtuse;

costa percurrent or excurrent as a smooth or weakly denticulate, occasionally decurrent awn, abaxial surface often papillose;

basal marginal cells usually quadrate;

distal laminal cells mostly short-rectangular, 8–10 µm wide, smooth, sinuose.

erect or weakly curved when dry, ovate-lanceolate to ovate-triangular strongly keeled distally, 1.3–1.9(–2.4) mm, 1-stratose, rarely with a few 2-stratose striae distally;

margins recurved to just before the apex, smooth, 2-stratose with 1-stratose patches;

apices acute;

costa excurrent as an often long, finely denticulate, straight or flexuose, and sometimes decurrent awn, often embracing part of the lamina, rarely percurrent, smooth;

basal marginal cells short-rectangular, usually with transverse walls thicker than longitudinal walls;

distal cells mainly ovate or short-rectangular, 7–10 µm wide, smooth, straight or slightly sinuose.

Sexual condition

autoicous.

autoicous.

Capsule

dark red or brown, short-cylindric, 0.7–1.3 mm;

exothecial cells usually quadrate, sometimes mixed with short-elongate or oblate cells, thin-walled, usually trigonous;

stomata present;

peristome patent or erect, often twisted, 350–600(–700) µm, red, densely papillose, usually perforated.

orange-brown or yellow-brown, cupulate, 0.6–0.9 mm;

exothecial often irregularly angular, short-elongate or isodiametric, thin-walled or slightly thickened, occasionally with small trigones;

rim red;

stomata present;

operculum mamillate;

peristome absent or rudimentary, consisting of basal segments, to 30 µm, yellowish or orange-red, papillose.

Spores

11–14(–19) µm, granulose or smooth.

8–10 µm, granulose or verruculose.

Schistidium apocarpum

Schistidium flaccidum

Phenology Capsules mature late spring to early summer. Capsules mature late spring to early summer.
Habitat Rocks in somewhat shaded habitats Rock in open to shaded habitats
Elevation low to moderate elevations (0-1500 m) (low to moderate elevations (0-4900 ft)) moderate to high elevations (1500-3200 m) (moderate to high elevations (4900-10500 ft))
Distribution
from FNA
AK; MI; NY; VT; WA; WI; AB; BC; NB; NF; NS; ON; QC; Greenland; Eurasia
[WildflowerSearch map]
from FNA
CO; ID; NV; AB; Eurasia; Africa; Atlantic Islands; Pacific Islands
Discussion

Although H. H. Blom (1996) considered Schistidium apocarpum to be restricted in the flora area to the eastern portions of North America, it is more widespread and scattered across the continent. The denticulate leaf margins, papillose abaxial costal surface, short-cylindrical capsules, and the thin-walled, often evenly quadrate exothecial cells are distinguishing characters. The long peristome teeth are also useful in identifying S. apocarpum. In good condition the peristome can only be confused with that of S. trichodon, which differs in the darker, often black color of its plants, the peristome teeth often forming a dome, and the cylindrical capsules that retain the columella.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Schistidium flaccidum is characterized by long and broad-based awns, light-colored capsules with red rims, the rudimentary peristome, and the mamillate rostrum. For comparison to the similar arctic taxon 10. S. cryptocarpum, see the discussion thereunder.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 211. FNA vol. 27, p. 216.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Schistidium Grimmiaceae > subfam. Grimmioideae > Schistidium
Sibling taxa
S. agassizii, S. atrichum, S. atrofuscum, S. boreale, S. cinclidodonteum, S. confertum, S. crassipilum, S. crassithecium, S. cryptocarpum, S. dupretii, S. flaccidum, S. flexipile, S. frigidum, S. frisvollianum, S. grandirete, S. heterophyllum, S. holmenianum, S. liliputanum, S. maritimum, S. occidentale, S. papillosum, S. pulchrum, S. rivulare, S. robustum, S. strictum, S. subjulaceum, S. tenerum, S. trichodon, S. venetum
S. agassizii, S. apocarpum, S. atrichum, S. atrofuscum, S. boreale, S. cinclidodonteum, S. confertum, S. crassipilum, S. crassithecium, S. cryptocarpum, S. dupretii, S. flexipile, S. frigidum, S. frisvollianum, S. grandirete, S. heterophyllum, S. holmenianum, S. liliputanum, S. maritimum, S. occidentale, S. papillosum, S. pulchrum, S. rivulare, S. robustum, S. strictum, S. subjulaceum, S. tenerum, S. trichodon, S. venetum
Synonyms Grimmia apocarpa, S. lancifolium, S. umbrosum Anictangium flaccidum, Grimmia flaccida
Name authority (Hedwig) Bruch & Schimper: Bryol. Europ. 3: 99. (1845) (De Notaris) Ochyra: Nova Hedwigia 48: 94. (1989)
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