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schistidium moss

compact grimmia, compact schistidium

Habit Plants in open tufts or mats, olivaceous to brownish (black), sometimes with yellowish tones. Plants in compact, often flattish cushions or tufts, olivaceous, brown, or yellow-brown, rarely nearly black.
Stems

1.2–12 cm, central strand weak or absent.

0.3–1.2(–2) cm, central strand narrow or absent.

Leaves

erect or curved, rarely (falcate-) secund when dry, ovate-lanceolate, sharply keeled distally, (1.3–)1.7–2.5(–3.2) mm, 1-stratose or rarely 2-stratose in striae distally;

margins usually recurved throughout or to just before the apex, usually denticulate distally, 1 or 2-stratose;

apices acute or sub-obtuse;

costa percurrent or excurrent as a smooth or weakly denticulate, occasionally decurrent awn, abaxial surface often papillose;

basal marginal cells usually quadrate;

distal laminal cells mostly short-rectangular, 8–10 µm wide, smooth, sinuose.

erect, sometimes slightly curved, imbricate when dry, ovate-lanceolate to ovate-triangular, obtusely keeled or concave proximally, sharply keeled distally, 0.8–1.8(–2) mm, 1-stratose or 2-stratose in striae distally;

margins recurved in medial sections, usually less recurved on one side of leaf, rarely plane, smooth, 2-stratose, occasionally 3-stratose;

apices usually acute, rarely sub-obtuse;

costa percurrent or excurrent as a short, spinulose-denticulate, often flexuose awn, smooth;

basal marginal cells quadrate or short-rectangular, transverse walls usually thicker than longitudinal walls, sometimes trigonous;

distal cells mostly isodiametric, rounded or angular, or short-rectangular, 5–8 µm wide, smooth, weakly sinuose.

Sexual condition

autoicous.

autoicous.

Capsule

dark red or brown, short-cylindric, 0.7–1.3 mm;

exothecial cells usually quadrate, sometimes mixed with short-elongate or oblate cells, thin-walled, usually trigonous;

stomata present;

peristome patent or erect, often twisted, 350–600(–700) µm, red, densely papillose, usually perforated.

yellow-brown or light reddish brown, short-cylindric or cupulate, rarely campanulate, rarely becoming finely striate with age, (0.5–)0.7–1 mm;

exothecial cells mostly short-elongate, mixed with patches of angular isodiametric cells, walls unevenly thickened, trigonous;

stomata present;

peristome patent to squarrose, sometimes twisted, 230–320 µm, orange or yellowish red, coarsely papillose, usually strongly perforated and with uneven margins.

Spores

11–14(–19) µm, granulose or smooth.

8–12 µm, smooth or granulose.

Schistidium apocarpum

Schistidium confertum

Phenology Capsules mature late spring to early summer. Capsules mature late spring to early summer.
Habitat Rocks in somewhat shaded habitats Rocks in somewhat shaded habitats
Elevation low to moderate elevations (0-1500 m) (low to moderate elevations (0-4900 ft)) moderate to high elevations (1000-3200 m) (moderate to high elevations (3300-10500 ft))
Distribution
from FNA
AK; MI; NY; VT; WA; WI; AB; BC; NB; NF; NS; ON; QC; Greenland; Eurasia
[WildflowerSearch map]
from FNA
AK; CA; CO; ID; MI; NV; UT; WA; WY; AB; BC; Greenland; Eurasia
Discussion

Although H. H. Blom (1996) considered Schistidium apocarpum to be restricted in the flora area to the eastern portions of North America, it is more widespread and scattered across the continent. The denticulate leaf margins, papillose abaxial costal surface, short-cylindrical capsules, and the thin-walled, often evenly quadrate exothecial cells are distinguishing characters. The long peristome teeth are also useful in identifying S. apocarpum. In good condition the peristome can only be confused with that of S. trichodon, which differs in the darker, often black color of its plants, the peristome teeth often forming a dome, and the cylindrical capsules that retain the columella.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Schistidium confertum is one of the smaller species of the genus. It can be confused with S. dupretii and S. frigidum. Schistidium dupretii is distinguished by its consistently striate, larger, darker, and more cylindrical capsules and its oblate basal marginal laminal cells versus the quadrate or longitudinally short- rectangular marginal cells of S. confertum. Also, S. dupretii lacks the thicker transverse walls that characterize the basal marginal cells of S. confertum. The leaf margins of S. confertum are more weakly recurved than in S. dupretii, and it is usually olivaceous, often with yellowish or brownish tones, whereas S. dupretii is generally evenly brown.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 211. FNA vol. 27, p. 214.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Schistidium Grimmiaceae > subfam. Grimmioideae > Schistidium
Sibling taxa
S. agassizii, S. atrichum, S. atrofuscum, S. boreale, S. cinclidodonteum, S. confertum, S. crassipilum, S. crassithecium, S. cryptocarpum, S. dupretii, S. flaccidum, S. flexipile, S. frigidum, S. frisvollianum, S. grandirete, S. heterophyllum, S. holmenianum, S. liliputanum, S. maritimum, S. occidentale, S. papillosum, S. pulchrum, S. rivulare, S. robustum, S. strictum, S. subjulaceum, S. tenerum, S. trichodon, S. venetum
S. agassizii, S. apocarpum, S. atrichum, S. atrofuscum, S. boreale, S. cinclidodonteum, S. crassipilum, S. crassithecium, S. cryptocarpum, S. dupretii, S. flaccidum, S. flexipile, S. frigidum, S. frisvollianum, S. grandirete, S. heterophyllum, S. holmenianum, S. liliputanum, S. maritimum, S. occidentale, S. papillosum, S. pulchrum, S. rivulare, S. robustum, S. strictum, S. subjulaceum, S. tenerum, S. trichodon, S. venetum
Synonyms Grimmia apocarpa, S. lancifolium, S. umbrosum Grimmia conferta
Name authority (Hedwig) Bruch & Schimper: Bryol. Europ. 3: 99. (1845) (Funck) Bruch & Schimper: Bryol. Europ. 3: 99. (1845)
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