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schistidium moss

Habit Plants in open tufts or mats, olivaceous to brownish (black), sometimes with yellowish tones. Plants in compact, often flat cushions or tufts, olivaceous, brown, or black.
Stems

1.2–12 cm, central strand weak or absent.

0.3–0.8 (–1.5) cm, central strand absent.

Leaves

erect or curved, rarely (falcate-) secund when dry, ovate-lanceolate, sharply keeled distally, (1.3–)1.7–2.5(–3.2) mm, 1-stratose or rarely 2-stratose in striae distally;

margins usually recurved throughout or to just before the apex, usually denticulate distally, 1 or 2-stratose;

apices acute or sub-obtuse;

costa percurrent or excurrent as a smooth or weakly denticulate, occasionally decurrent awn, abaxial surface often papillose;

basal marginal cells usually quadrate;

distal laminal cells mostly short-rectangular, 8–10 µm wide, smooth, sinuose.

erect, often curved towards stem when dry, ovate-ligulate to ovate-lanceolate, usually concave proximally, moderately keeled distally, 0.4–0.7(–0.8) mm, 1-stratose, sometimes with a few 2-stratose striae distally;

margins plane or recurved to just before the apex, smooth, 1-stratose or 2-stratose, rarely 3-stratose;

apices rounded or subacute;

costa sub-percurrent or percurrent, rarely excurrent as a tiny, weakly papillose awn, smooth;

basal marginal cells quadrate or short-rectangular;

distal laminal cells ovate or short-rectangular, 7–9 µm wide, smooth, sinuose.

Sexual condition

autoicous.

autoicous.

Capsule

dark red or brown, short-cylindric, 0.7–1.3 mm;

exothecial cells usually quadrate, sometimes mixed with short-elongate or oblate cells, thin-walled, usually trigonous;

stomata present;

peristome patent or erect, often twisted, 350–600(–700) µm, red, densely papillose, usually perforated.

dark red-brown or light brown, short-cylindric or cupulate, rarely slightly wider at the mouth, 0.4–0.6(–0.8) mm, occasionally finely ribbed when dry;

exothecial cells of various shapes, mainly elongate or isodiametric, walls thin or unevenly thickened, straight or curved, sometimes trigonous;

stomata absent;

peristome erect or patent, 100–200 µm, red-brown or red, papillose, weakly perforated.

Spores

11–14(–19) µm, granulose or smooth.

8–11 µm, smooth.

Schistidium apocarpum

Schistidium atrichum

Phenology Capsules mature late spring to early summer. Capsules mature late spring to early summer.
Habitat Rocks in somewhat shaded habitats Dry, often shaded mainly limestone rocks
Elevation low to moderate elevations (0-1500 m) (low to moderate elevations (0-4900 ft)) usually high elevations
Distribution
from FNA
AK; MI; NY; VT; WA; WI; AB; BC; NB; NF; NS; ON; QC; Greenland; Eurasia
[WildflowerSearch map]
from FNA
AK; CA; CO; ID; MT; OR; UT; WA; AB; BC
Discussion

Although H. H. Blom (1996) considered Schistidium apocarpum to be restricted in the flora area to the eastern portions of North America, it is more widespread and scattered across the continent. The denticulate leaf margins, papillose abaxial costal surface, short-cylindrical capsules, and the thin-walled, often evenly quadrate exothecial cells are distinguishing characters. The long peristome teeth are also useful in identifying S. apocarpum. In good condition the peristome can only be confused with that of S. trichodon, which differs in the darker, often black color of its plants, the peristome teeth often forming a dome, and the cylindrical capsules that retain the columella.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Schistidium atrichum is the smallest North American species in the genus. Along with the small size of its leaves, stems, and capsules, the usual lack of awns on vegetative leaves and its habit of forming small, often flat cushions help to distinguish it. S. Flowers (1973) noted strong similarities between S. atrichum and S. dupretii, but the larger size and the consistently ribbed capsules separate S. dupretii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 211. FNA vol. 27, p. 212.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Schistidium Grimmiaceae > subfam. Grimmioideae > Schistidium
Sibling taxa
S. agassizii, S. atrichum, S. atrofuscum, S. boreale, S. cinclidodonteum, S. confertum, S. crassipilum, S. crassithecium, S. cryptocarpum, S. dupretii, S. flaccidum, S. flexipile, S. frigidum, S. frisvollianum, S. grandirete, S. heterophyllum, S. holmenianum, S. liliputanum, S. maritimum, S. occidentale, S. papillosum, S. pulchrum, S. rivulare, S. robustum, S. strictum, S. subjulaceum, S. tenerum, S. trichodon, S. venetum
S. agassizii, S. apocarpum, S. atrofuscum, S. boreale, S. cinclidodonteum, S. confertum, S. crassipilum, S. crassithecium, S. cryptocarpum, S. dupretii, S. flaccidum, S. flexipile, S. frigidum, S. frisvollianum, S. grandirete, S. heterophyllum, S. holmenianum, S. liliputanum, S. maritimum, S. occidentale, S. papillosum, S. pulchrum, S. rivulare, S. robustum, S. strictum, S. subjulaceum, S. tenerum, S. trichodon, S. venetum
Synonyms Grimmia apocarpa, S. lancifolium, S. umbrosum Grimmia atricha
Name authority (Hedwig) Bruch & Schimper: Bryol. Europ. 3: 99. (1845) (Müller Hal. & Kindberg) W. A. Weber: Phytologia 33: 106. (1976)
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