Schistidium apocarpum |
Grimmiaceae subfam. grimmioideae |
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schistidium moss |
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Habit | Plants in open tufts or mats, olivaceous to brownish (black), sometimes with yellowish tones. | Plants acrocarpous. | ||||||||||||||||
Stems | 1.2–12 cm, central strand weak or absent. |
erect, central strand present or absent. |
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Leaves | erect or curved, rarely (falcate-) secund when dry, ovate-lanceolate, sharply keeled distally, (1.3–)1.7–2.5(–3.2) mm, 1-stratose or rarely 2-stratose in striae distally; margins usually recurved throughout or to just before the apex, usually denticulate distally, 1 or 2-stratose; apices acute or sub-obtuse; costa percurrent or excurrent as a smooth or weakly denticulate, occasionally decurrent awn, abaxial surface often papillose; basal marginal cells usually quadrate; distal laminal cells mostly short-rectangular, 8–10 µm wide, smooth, sinuose. |
erect or distally curved, rarely crisped, broadly oblong ovate, ovate-lanceolate, or narrowly lanceolate; margins plane, incurved or recurved, entire to occasionally denticulate distally, costa entire distally, percurrent to excurrent, smooth, in transverse section semi-terete, occasionally terete, elliptical or reniform, with 2 or occasionally with 3–6 adaxial cells near base, usually not markedly larger than abaxial cells, often excurrent as a long awn, awn smooth to toothed but not papillose; laminal cells smooth, mammillose, or papillose; basal cells quadrate to elongate, straight to sometimes sinuous, rarely sinuose-nodulose, thin- to thick-walled, without spiral thickenings, hyaline along insertion or concolorous with more distal basal cells; mid leaf cells mainly quadrate, to short rectangular, often sinuose but not sinuose-nodulose. |
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Seta | short to long, straight, arcuate, or sigmoid, smooth, one per perichaetium; vaginula with straight epidermal cells. |
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Sexual condition | autoicous. |
autoicous or dioicous. |
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Capsule | dark red or brown, short-cylindric, 0.7–1.3 mm; exothecial cells usually quadrate, sometimes mixed with short-elongate or oblate cells, thin-walled, usually trigonous; stomata present; peristome patent or erect, often twisted, 350–600(–700) µm, red, densely papillose, usually perforated. |
usually erect, rarely pendent, immersed to long-exserted, symmetric to ventricose, ovoid, obloid or cylindrical, sometimes globose, smooth or distinctly striate; stomata present or absent; annulus persistent or deciduous; operculum mammillate to long-rostrate; peristome absent basal membrane, distinctly thicker and trabeculate on the abaxial side, entire or split distally. |
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Calyptra | cucullate, mitrate, or mitrate-campanulate, smooth or plicate, not papillose, covering operculum to entire capsule. |
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Spores | 11–14(–19) µm, granulose or smooth. |
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Specialized | asexual reproduction absent or occasionally present as gemmae borne in axils of distal leaves or on leaf tips. |
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Schistidium apocarpum |
Grimmiaceae subfam. grimmioideae |
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Phenology | Capsules mature late spring to early summer. | |||||||||||||||||
Habitat | Rocks in somewhat shaded habitats | |||||||||||||||||
Elevation | low to moderate elevations (0-1500 m) (low to moderate elevations (0-4900 ft)) | |||||||||||||||||
Distribution |
AK; MI; NY; VT; WA; WI; AB; BC; NB; NF; NS; ON; QC; Greenland; Eurasia
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Worldwide |
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Discussion | Although H. H. Blom (1996) considered Schistidium apocarpum to be restricted in the flora area to the eastern portions of North America, it is more widespread and scattered across the continent. The denticulate leaf margins, papillose abaxial costal surface, short-cylindrical capsules, and the thin-walled, often evenly quadrate exothecial cells are distinguishing characters. The long peristome teeth are also useful in identifying S. apocarpum. In good condition the peristome can only be confused with that of S. trichodon, which differs in the darker, often black color of its plants, the peristome teeth often forming a dome, and the cylindrical capsules that retain the columella. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera ca. 8, species ca. 250 (5 genera, 81 species in the flora). Species of Grimmioideae are separated from those of the Racomitrioideae by features of the peristome, leaf cells that are straight to sinuose, and awns that are smooth to toothed. The Racomitrioideae have leaf cells that are strongly sinuose-nodulose, and awns that are sometimes smooth but are often papillose. The leaf attachments of the Racomitrioideae are brightly colored whereas those of Grimmioideae are hyaline or concolorous with the rest of the lamina (except Grimmia leibergii and G. attenuata which are yellow or orange). All Racomitrioideae have a straight or slightly arcuate seta that can be either smooth or papillose. Their capsules are smooth or almost so. In Grimmioideae the seta can be straight, arcuate, or sigmoid and are always smooth; capsules range from smooth to deeply plicate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27, p. 211. | FNA vol. 27, p. 205. | ||||||||||||||||
Parent taxa | Grimmiaceae > subfam. Grimmioideae > Schistidium | Grimmiaceae | ||||||||||||||||
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Synonyms | Grimmia apocarpa, S. lancifolium, S. umbrosum | |||||||||||||||||
Name authority | (Hedwig) Bruch & Schimper: Bryol. Europ. 3: 99. (1845) | Brotherus: in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. 215(I,3): 444. 1902 (as Grimmieae), | ||||||||||||||||
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