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Habit Plants medium-sized, green, yellowish, or with red secondary pigment. Plants medium-sized, with red or dark red secondary pigment, sometimes green.
Stem(s)

leaves ovate or ovate-triangular, gradually narrowed to apex, falcate or strongly so, sometimes straight, concave;

base not decurrent;

margins distinctly denticulate proximally, distally, or both;

apex acuminate, sometimes furrowed but not strongly so;

costa ending 3/5 to nearly entire leaf length;

alar region from margins to costa or nearly so;

marginal cells at widest part of leaf often differentiated from medial cells.

leaves oblong, ovate, or narrowly ovate, ± abruptly narrowed to apex, straight, concave;

base not or hardly decurrent;

margins entire or nearly so;

apex rounded- or acute-apiculate;

costa ending 4/5 to nearly entire leaf length;

alar region gradually passing into strongly incrassate cells near costa.

Sarmentypnum exannulatum

Sarmentypnum sarmentosum

Habitat Intermediately mineral-rich fens, around springs, late snow beds, submerged in lakes Intermediately mineral-rich fens, around springs, late snow beds, submerged in lakes
Elevation low to high elevations (0-4200 m) (low to high elevations (0-13800 ft)) low to high elevations (0-4000 m) (low to high elevations (0-13100 ft))
Distribution
from FNA
AK; CA; CO; ID; MA; MD; ME; MI; MN; MT; NH; NV; NY; OR; PA; UT; WA; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Pacific Islands (New Zealand); Greenland; Africa; Eurasia; Australia
from FNA
AK; CO; MT; NH; NY; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; YT; South America; Pacific Islands (New Zealand, Papua New Guinea); Australia; Greenland; e Africa; Eurasia; Antarctica
Discussion

Sarmentypnum exannulatum is one of the most frequent wetland species, with leaves that narrow gradually to the apex and are gradually curved throughout. The marginal laminal cells in the widest part of the stem leaves are often distinctly rectangular, distinctly widened, or both; this feature is never as pronounced in other species of Sarmentypnum as in S. exannulatum. However, because this feature is rather variable it should be used with caution when identifying material. Warnstorfia fluitans and W. pseudostraminea differ from S. exannulatum in being autoicous, having mainly narrowly triangular to lanceolate pseudoparaphyllia, hardly ever with pure red pigments, and with more weakly differentiated alar regions. The alar regions in W. fluitans usually reach less far distally in the leaf than in S. exannulatum, whereas those in W. pseudostraminea often form ovate regions together with the supra-alar cells. Phenotypes of S. exannulatum, found in springs, with inflated alar cells in more or less one row along the leaf base have been called W. exannulata var. purpurascens. However, transitions between these phenotypes and those from other habitats exist, and if the different phenotypes are cultivated together, the resulting plants cannot be separated (based on north European material).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sarmentypnum sarmentosum is distinguished by the oblong or ovate stem leaves suddenly narrowed to rounded- or acute-apiculate apices. The apiculus is most distinct in young leaves and is rarely completely absent, but may be eroded in older leaves. The species could be confused with species of Calliergon, which are larger, have broader leaves, and never have a clear red color, or with Straminergon stramineum, which is never red and lacks an apiculus. Straminergon stramineum is more sparsely branched (sometimes unbranched) than Sarmentypnum sarmentosum, and the alar regions are ovate or broadly ovate in Straminergon, transversely triangular in S. sarmentosum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 400. FNA vol. 28, p. 403.
Parent taxa Calliergonaceae > Sarmentypnum Calliergonaceae > Sarmentypnum
Sibling taxa
S. pseudosarmentosum, S. sarmentosum, S. trichophyllum, S. tundrae
S. exannulatum, S. pseudosarmentosum, S. trichophyllum, S. tundrae
Synonyms Hypnum exannulatum, Drepanocladus exannulatus, D. exannulatus var. purpurascens, D. exannulatus var. rotae, Warnstorfia exannulata, W. exannulata var. purpurascens Hypnum sarmentosum, Calliergon sarmentosum, C. sarmentosum var. beringianum, C. sarmentosum var. fallaciosum, C. sarmentosum var. fontinaloides, Warnstorfia sarmentosa
Name authority (Schimper) Hedenas: J. Hattori Bot. Lab. 100: 132. (2006) (Wahlenberg) Tuomikoski & T. J. Koponen: Ann. Bot. Fenn. 16: 223. (1979)
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