Sandbergia whitedii |
Sandbergia |
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fissurewort, Whited's fissurewort, Whited's halimolobos, Whited's halimolobos fissurewort |
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Habit | Perennials; densely and uniformly pubescent, trichomes shortly stalked, minute, Y-shaped or cruciform, mixed with simple or 1 or more ray-forked ones. | Biennials or perennials; (caudex simple or branched, covered with persistent leaf remains); not scapose; pubescent throughout, trichomes shortly stalked or subsessile, cruciform, Y-shaped, or forked. | ||||
Stems | simple or few from caudex, erect, branched (several) distally, (1.5–)2.5–5.5(–6.5) dm, densely pubescent throughout. |
erect or decumbent, unbranched or branched distally. |
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Leaves | basal and cauline; petiolate or sessile; basal usually rosulate, petiolate, blade margins entire, dentate, or lyrate-pinnatifid, (apex obtuse to acute); cauline sessile, blade (base attenuate, not auriculate), margins entire, subentire, dentate, or pinnatifid. |
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Basal leaves | often rosulate; petiole (1–)2–5(–7) cm; blade spatulate to linear-oblanceolate, (1–)1.7–5 cm × (3–)4–9(–12) mm, margins entire or dentate, surfaces densely pubescent. |
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Cauline leaves | 3–7 (widely spaced); blade linear-oblanceolate, 2–7 cm × 1–5(–8) mm, margins usually entire, rarely minutely sparsely denticulate, apex obtuse, surfaces densely pubescent. |
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Racemes | (corymbose, several-flowered), considerably elongated in fruit. |
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Flowers | sepals 2.5–3 × 1–1.5 mm; petals (4–)5–6.5(–7.5) × (1.2–)1.5–2(–2.2) mm; filaments 1.5–3 mm; anthers oblong, 0.6–0.8 mm. |
sepals (erect), oblong; petals white, oblanceolate-spatulate, (longer than sepals, claw obscurely differentiated from blade, apex rounded); stamens slightly tetradynamous; filaments not dilated basally, (slender); anthers ovate or oblong, (apex obtuse); nectar glands confluent, subtending bases of stamens. |
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Fruiting pedicels | (5–)8–14(–17) mm, densely pubescent. |
ascending to subdivaricate, (straight), slender, (terete). |
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Fruits | suberect, (not appressed to rachis), straight, slightly torulose, subterete, (1–)1.3–1.8(–2) cm × 0.8–1 mm; valves each with obscure midvein on proximal 1/2, densely pubescent; ovules 20–30 per ovary; style 0.2–0.4 mm. |
subsessile or shortly stipitate (gynophore less than 1 mm), linear, slightly to strongly torulose, subterete to strongly latiseptate; valves each without midvein or with obscure one on proximal 1/2, sparsely to densely pubescent or glabrescent; replum rounded; septum complete; ovules 12–30 per ovary; (style obsolete or distinct); stigma capitate. |
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Seeds | 1–1.2 × 0.6–0.7 mm. |
uniseriate, plump, not winged, oblong; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons incumbent. |
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x | = 7. |
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Sandbergia whitedii |
Sandbergia |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Dry sagebrush scabland, gravelly hillsides, basaltic talus, dry sandy slopes, alpine meadows, cliffs, ridge crests | |||||
Elevation | 500-1200 m (1600-3900 ft) | |||||
Distribution |
WA; BC
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nw North America |
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Discussion | In Washington state, Sandbergia whitedii appears to be restricted to Chelan, Douglas, Grant, Kittitas, Lincoln, and Okanogan counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 2 (2 in the flora). Although both species of Sandbergia were placed by R. C. Rollins (1993) in Halimolobos, the two genera are not closely related. Sandbergia is most closely related to Boechera, whereas Halimolobos is sister to Mancoa and Sphaerocardamum Schauer in the tribe Halimolobeae. For an account of the generic boundaries of Sandbergia, see under 60. Halimolobos. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 418. | FNA vol. 7, p. 417. | ||||
Parent taxa | Brassicaceae > tribe Boechereae > Sandbergia | Brassicaceae > tribe Boechereae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Arabis whitedii, Halimolobos whitedii | |||||
Name authority | (Piper) Greene: Leafl. Bot. Observ. Crit. 2: 137. (1911) | Greene: Leafl. Bot. Observ. Crit. 2: 136. (1911) | ||||
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