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Habit Biennials or perennials; (caudex simple or branched, covered with persistent leaf remains); not scapose; pubescent throughout, trichomes shortly stalked or subsessile, cruciform, Y-shaped, or forked. Perennials or, rarely, biennials; eglandular.
Stems

erect or decumbent, unbranched or branched distally.

Leaves

basal and cauline;

petiolate or sessile;

basal usually rosulate, petiolate, blade margins entire, dentate, or lyrate-pinnatifid, (apex obtuse to acute);

cauline sessile, blade (base attenuate, not auriculate), margins entire, subentire, dentate, or pinnatifid.

Cauline leaves

(sometimes absent);

petiolate, subsessile, or sessile;

blade base auriculate or not, margins usually entire or dentate, rarely pinnatifid.

Trichomes

often short-stalked, sessile, or subsessile, usually forked or dendritic, rarely malpighiaceous, sometimes simple or absent.

Racemes

(corymbose, several-flowered), considerably elongated in fruit.

ebracteate, often elongated in fruit.

Flowers

sepals (erect), oblong;

petals white, oblanceolate-spatulate, (longer than sepals, claw obscurely differentiated from blade, apex rounded);

stamens slightly tetradynamous;

filaments not dilated basally, (slender);

anthers ovate or oblong, (apex obtuse);

nectar glands confluent, subtending bases of stamens.

actinomorphic;

sepals erect, ascending, or spreading, lateral pair usually not saccate basally;

petals white, yellowish, pink, lavender, or purple, claw present, often indistinct;

filaments unappendaged, not winged;

pollen 3-colpate.

Fruiting pedicels

ascending to subdivaricate, (straight), slender, (terete).

Fruits

subsessile or shortly stipitate (gynophore less than 1 mm), linear, slightly to strongly torulose, subterete to strongly latiseptate;

valves each without midvein or with obscure one on proximal 1/2, sparsely to densely pubescent or glabrescent;

replum rounded;

septum complete;

ovules 12–30 per ovary; (style obsolete or distinct);

stigma capitate.

usually siliques, rarely silicles, dehiscent, unsegmented, usually latiseptate, rarely terete or slightly angustiseptate;

ovules [2–]4–250 per ovary;

style usually distinct, rarely obsolete;

stigma usually entire, rarely 2-lobed.

Seeds

uniseriate, plump, not winged, oblong;

seed coat (minutely reticulate), not mucilaginous when wetted;

cotyledons incumbent.

biseriate, sub-biseriate, uniseriate, or, rarely, aseriate;

cotyledons accumbent or incumbent.

x

= 7.

Sandbergia

Brassicaceae tribe Boechereae

Distribution
nw North America
[BONAP county map]
North America; Asia (Russian Far East)
Discussion

Species 2 (2 in the flora).

Although both species of Sandbergia were placed by R. C. Rollins (1993) in Halimolobos, the two genera are not closely related. Sandbergia is most closely related to Boechera, whereas Halimolobos is sister to Mancoa and Sphaerocardamum Schauer in the tribe Halimolobeae. For an account of the generic boundaries of Sandbergia, see under 60. Halimolobos.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 7, species 119 (7 genera, 117 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf blade margins usually entire, rarely denticulate; fruits slightly torulose.
S. whitedii
1. Cauline leaf blade margins coarsely dentate to pinnatifid; fruits strongly torulose.
S. perplexa
Source FNA vol. 7, p. 417. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 347.
Parent taxa Brassicaceae > tribe Boechereae Brassicaceae
Subordinate taxa
S. perplexa, S. whitedii
Name authority Greene: Leafl. Bot. Observ. Crit. 2: 136. (1911) Al-Shehbaz: Pl. Syst Evol. 259: 111. (2006)
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