Salix viminalis |
Salix sitchensis |
|
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basket willow, common osier, osier, osier willow, silky osier |
Coulter willow, Sitka willow |
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Habit | Shrubs or trees, 1–8 m, (sometimes forming clones by stem fragmentation). | |
Stems | branches yellow-brown, gray-brown, or yellowish, not glaucous, glabrous or puberulent; branchlets yellow-brown or yellowish (sometimes color obscured by hairs), glabrous, densely to sparsely villous, velvety, or puberulent. |
branches (sometimes highly brittle at base), yellow-brown or red-brown, not glaucous, glabrous or pilose; branchlets yellow-brown, gray-brown, or red-brown, densely short-silky, velvety, or villous, (buds caprea-type or intermediate). |
Leaves | stipules (not adnate to petioles), rudimentary or absent on early ones, (late ones sometimes brownish, linear, 5.4–10.4 mm), apex acuminate; petiole shallowly grooved adaxially, 4–13 mm, villous, puberulent, or velvety adaxially; largest medial blade linear, lorate, narrowly oblong, or narrowly elliptic, 53–130 × 5–33 mm, base cuneate, margins strongly revolute, sinuate or apparently entire, (glands epilaminal), apex acuminate, acute, or convex, abaxial surface apparently glaucous (obscured by hairs), densely short-silky, woolly, or tomentose, (midribs prominent, yellowish, and hairy), hairs appressed, spreading or erect, straight or wavy, adaxial dull or slightly glossy, sparsely or moderately densely pubescent, hairs gray; proximal blade margins entire; juvenile blade yellowish green, very densely tomentose or short-silky abaxially, hairs white. |
stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute; petiole convex to flat, or shallowly grooved adaxially, 3–13(–16) mm, tomentose or velvety adaxially; largest medial blade elliptic, narrowly oblanceolate, oblanceolate, or obovate, 31–70–120 × 17–48 mm, 2.1–3.1–4 times as long as wide, base cuneate or convex, margins slightly revolute or flat, strongly revolute proximally, entire, irregularly serrate, or sinuate, (glands submarginal or epilaminal), apex acuminate or convex, abaxial surface not evidently glaucous, (obscured by hairs), densely short-silky, woolly, or silky-woolly, hairs straight, wavy, or curved, adaxial slightly glossy (sometimes dull and glaucous), pilose or moderately densely short-silky; proximal blade margins entire or shallowly serrulate; juvenile blade green, densely long-silky or woolly abaxially, (sparsely silky-tomentose adaxially), hairs white. |
Staminate flowers | adaxial nectary narrowly oblong or oblong, 0.6–1.5 mm; filaments distinct; anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.6–0.8 mm. |
adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.4–1.3 mm; stamens 1; filaments distinct, glabrous; anthers purple turning yellow, shortly cylindrical, 0.5–0.7 mm. |
Pistillate flowers | adaxial nectary narrowly oblong or oblong, 0.9–1.4 mm; ovary pyriform, beak gradually tapering to styles; ovules 12–18 per ovary; styles 0.6–1.8 mm. |
adaxial nectary square, ovate, or flask-shaped, 0.5–0.9 mm, shorter to longer than stipe; stipe 0.4–1.4 mm; ovary ovoid to pyriform, long- or short-silky or villous, beak sometimes slightly bulged below styles; ovules 14–20 per ovary; styles 0.4–0.8 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, 0.16–0.28–0.4 mm. |
Capsules | 4–6 mm. |
3.5–5.6 mm. |
Catkins | flowering just before or as leaves emerge; staminate stout, 24–48 mm, flowering branchlet 0–2 mm; pistillate densely flowered, 23–55 mm, flowering branchlet 0–6 mm; floral bract brown or tawny, 1.6–2.2 mm, apex convex or rounded, abaxially hairy, hairs straight. |
flowering just before or as leaves emerge; staminate slender or stout, (17–)22–54 × 8–15 mm, flowering branchlet 1–9 mm; pistillate moderately densely flowered, slender to stout, 25–73(–115 in fruit) × 5–15 mm, flowering branchlet 1–20 mm; floral bract tawny to dark brown, 1.4–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight or wavy. |
2n | = 38. |
= 38. |
Salix viminalis |
Salix sitchensis |
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Phenology | Flowering Apr-early May. | Flowering early Apr-mid Jun (Mar in California). |
Habitat | Sandy, open woods, cobble rivershores, lake margins, and roadsides | Tidal swamps and marshes, coastal fog belts and headlands, sand dunes, springs, gravelly streambeds and deltas, glacial moraines, avalanche tracks, dry canyons, clearings and edges of forests, shade tolerant |
Elevation | 0-300 m (0-1000 ft) | 0-1800 m (0-5900 ft) |
Distribution |
CT; IA; IN; MA; ME; NJ; NY; OH; RI; VT; NB; NF; NS; ON; PE; QC; Europe [Introduced in North America] |
AK; CA; ID; MT; OR; WA; AB; BC
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Discussion | Salix ×smithiana (S. caprea × S. viminalis) is distinguished from S. viminalis by having leaf blades usually broad, 2.8–4.9(–6.4) times as long as wide, stipes 0.9–2 mm, ovaries short-silky, branches ± brittle at base, and petioles flat to convex adaxially; S. viminalis has leaf blades usually very narrow, 4.7–13.7 times as long as wide, stipes 0.1–0.5 mm, ovaries long-silky, branches flexible at base, and petioles shallowly grooved adaxially. See Salix ×smithiana [p. 132] and 86. S. pellita for further comparative descriptions. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Ovary hairiness in some Salix sitchensis populations varies from uniformly hairy to glabrescent, with intermediates with patchy or streaky hairiness. All three variations can occur together and do not seem to indicate hybridization. Both Salix sitchensis and S. scouleriana have similar variants with leaves having very densely curly hairs on abaxial surfaces [S. sitchensis forma coulteri (Andersson) Jepson and S. scouleriana forma poikila (C. K. Schneider) C. K. Schneider]. Plants resembling S. drummondiana but with similar indumentum probably are hybrids with S. alaxensis (see 84. S. drummondiana). The coulteri taxon resembles S. delnortensis in having stipules with adaxial surfaces glabrous and very sparsely glandular toward the base, densely hairy abaxially, and with gland-dotted margins; its branchlets have wavy to crinkly hairs. The possible hybrid origin of S. delnortensis needs study (R. D. Dorn 2000). Hybrids: Salix sitchensis forms natural hybrids with S. alaxensis var. longistylis and S. melanopsis. Hybridization with S. geyeriana reported by J. K. Henry (1915) is not based on convincing specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 149. | FNA vol. 7, p. 159. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Viminella | Salicaceae > Salix > subg. Vetrix > sect. Sitchenses |
Sibling taxa | ||
Synonyms | S. coulteri, S. sitchensis var. parvifolia | |
Name authority | Linnaeus: Sp. Pl. 2: 1021. (1753) | Sanson ex Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 162. (1832) |
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