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rock willow

skeleton-leaf willow

Habit Plants 0.2–1.5 m. Stems erect; branches brownish or red-brown, (dull), glabrous, long-silky, or villous to glabrescent; branchlets yellow-brown or gray-brown, long-silky, pilose, or moderately densely villous. Plants 0.01–0.07 m, (dwarf), forming clonal mats by rhizomes.
Stems

trailing;

branches red-brown or yellow-brown, glabrous;

branchlets red-brown, glabrous.

Leaves

stipules absent or rudimentary;

petiole (shallowly to deeply grooved adaxially), 2–8 mm, (with 2 spherical glands distally, dark brown, sometimes basilaminar, sparsely pubescent or glabrous adaxially);

largest medial blade hypostomatous, (veins strongly impressed-reticulate), broadly elliptic, subcircular, or obovate, 18–67 × 10–40 mm, 1.1–2.3 times as long as wide, base rounded, convex, or subcordate, margins strongly revolute, crenate or subentire, apex rounded, convex, retuse, or toothed, abaxial surface sparsely to densely villous or long-silky, veins often with long, straight hairs, adaxial slightly glossy, glabrous or sparsely long-silky;

proximal blade margins entire or crenate;

juvenile blade (yellowish green), abaxially very densely long-silky.

(prominently marcescent, becoming skeletonized);

stipules absent or rudimentary;

petiole (deeply grooved or convex to flat adaxially), 1.2–3.2(–4.8) mm, (sparsely pubescent adaxially);

largest medial blade amphistomatous, (2 pairs secondary veins arising at or close to base, arcing toward apex), elliptic, broadly elliptic, obovate, subcircular, or circular, 7–15 × 3–11 mm, 1.1–2.5(–3.5) times as long as wide, base convex or cuneate, margins flat, entire, sometimes ciliate, apex convex, rounded, or retuse, abaxial surface not glaucous, glabrous, midrib sometimes pilose, hairs long, straight, wavy, or crinkled, adaxial highly glossy, glabrous;

proximal blade margins entire;

juvenile blade (green), glabrous abaxially or ciliate.

Staminate flowers

abaxial nectary 0.6–0.8 mm, adaxial nectary narrowly oblong, 0.5–1.2 mm, nectaries connate and shallowly cup-shaped, or distinct;

filaments distinct, hairy on proximal 1/2;

anthers ellipsoid or globose, 0.3–0.5 mm.

abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.4–1.1 mm;

filaments distinct or connate less than 1/2 their lengths, glabrous;

anthers ellipsoid or obovoid, 0.3–0.5 mm.

Pistillate flowers

abaxial nectary (0–)0.8–0.9 mm, adaxial nectary oblong, ovate, or narrowly oblong to almost filiform, 0.7–1.4 mm, shorter to longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0.4–1.2 mm;

ovary pyriform or obnapiform, densely short-silky, hairs cylindrical, beak abruptly tapering to styles;

ovules 13–15 per ovary;

styles connate 1/2 their lengths to almost distinct, 0.2–0.4 mm;

stigmas flat, abaxially non-papillate with rounded tip, slenderly cylindrical, or 2 plump lobes, 0.2–0.28–0.36 mm.

abaxial nectary absent, adaxial nectary oblong or narrowly oblong, 0.4–1.6 mm, shorter than or equal to stipe;

stipe 0.4–1.4 mm;

ovary pyriform, sparsely to moderately densely short-silky or villous, at least on beaks, hairs ribbonlike, beak slightly bulged below styles;

ovules 12 per ovary;

styles connate or slightly distinct distally, 0.3–1 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly to broadly cylindrical, 0.16–0.34–0.52 mm.

Capsules

3–5 mm.

2.9–4.8 mm.

Catkins

staminate 13–48 × 4–6.5(–8) mm, flowering branchlet 3–31(–50) mm; pistillate densely flowered, slender or stout, 18–56 × 4–10 mm, flowering branchlet 3–27(–40) mm;

floral bract tawny, 0.8–1.6 mm, apex rounded, entire, abaxially densely hairy, hairs straight.

staminate 10–35 × 6–10 mm, flowering branchlet 1–11 mm; pistillate moderately densely flowered, stout or subglobose, 12–38 × 5–11 mm, flowering branchlet 3–14 mm;

floral bract brown, black, or bicolor, 1–1.3 mm, apex rounded, entire, abaxially sparsely hairy, hairs straight or wavy.

2n

= 38.

= 38.

Salix vestita

Salix phlebophylla

Phenology Flowering mid Jun-late Jul. Flowering Jun–Jul.
Habitat Moist to dry open forests and rocky streamsides, in upper montane and subalpine zones, rarely alpine Arctic-alpine, dry Dryas-lichen tundra, polygonal tundra with stone stripes and dry raised centers, scree and colluvial slopes, grass-sedge tussock tundra, sedge meadows in drainage ways, dwarf birch thickets, granitic and sandstone substrates
Elevation 0-2400 m (0-7900 ft) 0-2100 m (0-6900 ft)
Distribution
from FNA
MT; OR; WA; AB; BC; MB; NL; NS; NU; ON; QC; Asia (China [Xinjiang], Mongolia, Russia, e, c Siberia)
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; YT; e Asia (Chukotka, Russian Far East, arctic, e Siberia)
[BONAP county map]
Discussion

Salix vestita is an ancient amphiberingian species. Its distribution includes a series of isolated, disjunct populations in Central Siberia, the northern Rocky Mountains, the west coast of Hudson Bay, and the northeastern arctic and subarctic. Occurrence in Nunavut is on Akpatok Island in Ungava Bay and on the Belcher Islands in Hudson Bay. It may be extirpated in Washington.

The flowering and vegetative branchlets sometimes have relatively short internodes. In subsequent years, branches have the appearance of short shoots similar to those in Alnus. Short shoots do not appear on all branches or in all years. The formation of short shoots may be related to adverse growing conditions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybrids:

Salix phlebophylla forms natural hybrids with S. arctica, S. fuscescens, and S. rotundifolia.

Salix phlebophylla × S. rotundifolia has hairy ovaries, some skeletonized leaves, a compact growth form that may lack rhizomes, and catkins with more than 15 flowers. It occurs on the Alaska arctic slope, outside the range of S. polaris.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 64. FNA vol. 7, p. 74.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Chamaetia > sect. Myrtosalix
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. leiolepis, S. vestita subsp. leiolepis, S. vestita var. psilophylla
Name authority Pursh: Fl. Amer. Sept. 2: 610. (1813) Andersson: Proc. Amer. Acad. Arts 4: 72. (1858)
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