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rock willow

gray willow, large gray or gray willow, large gray willow

Habit Plants 0.2–1.5 m. Stems erect; branches brownish or red-brown, (dull), glabrous, long-silky, or villous to glabrescent; branchlets yellow-brown or gray-brown, long-silky, pilose, or moderately densely villous. Shrubs, 3–7 m. Stems: branches brownish, not glaucous, pilose, villous, or tomentose to glabrescent, (peeled wood with striae to 62 mm); branchlets yellow-brown, pilose, velvety, or densely villous.
Leaves

stipules absent or rudimentary;

petiole (shallowly to deeply grooved adaxially), 2–8 mm, (with 2 spherical glands distally, dark brown, sometimes basilaminar, sparsely pubescent or glabrous adaxially);

largest medial blade hypostomatous, (veins strongly impressed-reticulate), broadly elliptic, subcircular, or obovate, 18–67 × 10–40 mm, 1.1–2.3 times as long as wide, base rounded, convex, or subcordate, margins strongly revolute, crenate or subentire, apex rounded, convex, retuse, or toothed, abaxial surface sparsely to densely villous or long-silky, veins often with long, straight hairs, adaxial slightly glossy, glabrous or sparsely long-silky;

proximal blade margins entire or crenate;

juvenile blade (yellowish green), abaxially very densely long-silky.

stipules rudimentary or foliaceous on early ones, foliaceous on late ones, apex acute or rounded;

petiole convex to flat adaxially, 4–15 mm, tomentose adaxially;

largest medial blade elliptic, broadly elliptic, oblanceolate, or obovate, 65–105 × 22–52 mm, 2–3 times as long as wide, base convex or cuneate, margins slightly revolute, entire, crenate, or sinuate, (glands submarginal), apex acuminate or convex, abaxial surface glaucous, tomentose, hairs erect or spreading, curly, adaxial dull or slightly glossy, pubescent or tomentose;

proximal blade margins entire;

juvenile blade yellowish green, sparsely to densely tomentose abaxially, hairs white.

Staminate flowers

abaxial nectary 0.6–0.8 mm, adaxial nectary narrowly oblong, 0.5–1.2 mm, nectaries connate and shallowly cup-shaped, or distinct;

filaments distinct, hairy on proximal 1/2;

anthers ellipsoid or globose, 0.3–0.5 mm.

adaxial nectary oblong or ovate, 0.5–1 mm;

filaments distinct, glabrous or hairy basally;

anthers yellow or purple turning yellow, ellipsoid or shortly cylindrical, 0.7–1 mm.

Pistillate flowers

abaxial nectary (0–)0.8–0.9 mm, adaxial nectary oblong, ovate, or narrowly oblong to almost filiform, 0.7–1.4 mm, shorter to longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0.4–1.2 mm;

ovary pyriform or obnapiform, densely short-silky, hairs cylindrical, beak abruptly tapering to styles;

ovules 13–15 per ovary;

styles connate 1/2 their lengths to almost distinct, 0.2–0.4 mm;

stigmas flat, abaxially non-papillate with rounded tip, slenderly cylindrical, or 2 plump lobes, 0.2–0.28–0.36 mm.

adaxial nectary oblong or square, 0.4–1 mm, shorter than stipe;

stipe 1.2–2.7 mm;

ovary pyriform, long-silky, beak slightly bulged below styles;

ovules 12 per ovary;

styles 0.2–0.5 mm;

stigmas slenderly or broadly cylindrical, 0.3–0.6 mm.

Capsules

3–5 mm.

5–5.6 mm.

Catkins

staminate 13–48 × 4–6.5(–8) mm, flowering branchlet 3–31(–50) mm; pistillate densely flowered, slender or stout, 18–56 × 4–10 mm, flowering branchlet 3–27(–40) mm;

floral bract tawny, 0.8–1.6 mm, apex rounded, entire, abaxially densely hairy, hairs straight.

flowering before leaves emerge; staminate stout or subglobose, 26–39 × 12–26 mm, flowering branchlet 0–5 mm; pistillate densely flowered, stout or subglobose, 27–54(–75 in fruit) × 4–15 mm, flowering branchlet 1–5(–10) mm;

floral bract dark brown, black, or bicolor, 2–3 mm, apex acute or convex, abaxially hairy, hairs straight.

2n

= 38.

= 76.

Salix vestita

Salix cinerea

Phenology Flowering mid Jun-late Jul. Flowering mid Mar-late May.
Habitat Moist to dry open forests and rocky streamsides, in upper montane and subalpine zones, rarely alpine Stream shores, mesic woodlands, gravelly or sandy beaches, waste ground
Elevation 0-2400 m (0-7900 ft) 0-700 m (0-2300 ft)
Distribution
from FNA
MT; OR; WA; AB; BC; MB; NL; NS; NU; ON; QC; Asia (China [Xinjiang], Mongolia, Russia, e, c Siberia)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; GA; IA; KY; LA; MA; MD; MI; MO; NC; NJ; NY; OH; PA; RI; SC; SD; TN; UT; VA; WI; WV; ON; Eurasia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix vestita is an ancient amphiberingian species. Its distribution includes a series of isolated, disjunct populations in Central Siberia, the northern Rocky Mountains, the west coast of Hudson Bay, and the northeastern arctic and subarctic. Occurrence in Nunavut is on Akpatok Island in Ungava Bay and on the Belcher Islands in Hudson Bay. It may be extirpated in Washington.

The flowering and vegetative branchlets sometimes have relatively short internodes. In subsequent years, branches have the appearance of short shoots similar to those in Alnus. Short shoots do not appear on all branches or in all years. The formation of short shoots may be related to adverse growing conditions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The Ohio occurrence of Salix cinerea is based on information from T. Cooperrider (pers. comm.).

Salix cinerea and S. atrocinerea are very closely related. Their occurrence in the flora area, as naturalized introductions, is not well understood, probably because they usually are introduced under the name S. caprea, and that species often is not treated in North American floristic literature (e.g., C. K. Schneider 1921; M. L. Fernald 1950). They probably are introductions of long-standing brought to the New World for their value as ornamentals and bee-plants. Salix atrocinerea was first documented in the southeastern United States (G. W. Argus 1986) after plants with ferruginous hairs and prominently striate wood were found in North Carolina; since that time, it has been found in other states and provinces. In the northeastern states, S. atrocinerea and S. cinerea are thought to be invasive species. The species do reproduce by seed and hundreds of seedlings were observed in a drained reservoir (A. Zinovjev, pers. comm.) and on sandy pond shores (T. Rawinski, pers. comm.), where they are thought to compete with native species.

The presence of long, prominent, striae on the peeled wood of 4–5 year old branches is commonly used in European literature (K. H. Rechinger 1993; A. K. Skvortsov 1999) to separate Salix cinerea and S. atrocinerea from S. caprea etc., in which the wood is smooth or with fewer, shorter striae. In the flora area, long striae also occur in S. bebbiana, S. discolor, and S. humilis, but usually they are not as long as or as prominent in S. cinerea and S. atrocinerea. Some floras (e.g., F. Martini and P. Paiero 1988) use the relative prominence of striae to separate S. cinerea and S. atrocinerea, but their separation remains difficult. The presence of ferruginous hairs on the leaves of S. atrocinerea is the best diagnostic characteristic, but they are not always present or easily observed. For a comparison of these species, see the key to species under subg. Vetrix. For further discussion of morphologies, see Salix ×smithiana Willdenow [p. 132] and 76. S. discolor.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 64. FNA vol. 7, p. 132.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Vetrix > sect. Cinerella
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. leiolepis, S. vestita subsp. leiolepis, S. vestita var. psilophylla
Name authority Pursh: Fl. Amer. Sept. 2: 610. (1813) Linnaeus: Sp. Pl. 2: 1021. (1753)
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