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rock willow

Athabasca willow

Habit Plants 0.2–1.5 m. Stems erect; branches brownish or red-brown, (dull), glabrous, long-silky, or villous to glabrescent; branchlets yellow-brown or gray-brown, long-silky, pilose, or moderately densely villous. Plants 0.6–1.3 m, not clonal.
Stems

erect;

branches gray-brown, hairy;

branchlets red-brown, sparsely or moderately densely pubescent, (buds alba-type).

Leaves

stipules absent or rudimentary;

petiole (shallowly to deeply grooved adaxially), 2–8 mm, (with 2 spherical glands distally, dark brown, sometimes basilaminar, sparsely pubescent or glabrous adaxially);

largest medial blade hypostomatous, (veins strongly impressed-reticulate), broadly elliptic, subcircular, or obovate, 18–67 × 10–40 mm, 1.1–2.3 times as long as wide, base rounded, convex, or subcordate, margins strongly revolute, crenate or subentire, apex rounded, convex, retuse, or toothed, abaxial surface sparsely to densely villous or long-silky, veins often with long, straight hairs, adaxial slightly glossy, glabrous or sparsely long-silky;

proximal blade margins entire or crenate;

juvenile blade (yellowish green), abaxially very densely long-silky.

stipules absent or rudimentary on early ones, usually rudimentary, rarely foliaceous, on late ones;

petiole (shallowly grooved adaxially), 3–10 mm, (puberulent or villous);

largest medial blade oblong, narrowly elliptic, elliptic, oblanceolate, or obovate, 17–50 × 8–18 mm, 1.9–3.2 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex acuminate or convex, abaxial surface glabrescent or sparsely silky, hairs appressed or somewhat spreading, (usually white, sometimes also ferruginous), straight or wavy, adaxial dull or slightly glossy, glabrous, glabrescent, pilose, or sparsely long-silky along midribs and margin, (hairs usually white, sometimes also ferruginous, appressed);

proximal blade margins entire;

juvenile blade sparsely to moderately densely villous or long-silky abaxially (hairs usually white, sometimes also ferruginous).

Staminate flowers

abaxial nectary 0.6–0.8 mm, adaxial nectary narrowly oblong, 0.5–1.2 mm, nectaries connate and shallowly cup-shaped, or distinct;

filaments distinct, hairy on proximal 1/2;

anthers ellipsoid or globose, 0.3–0.5 mm.

abaxial nectary (0–)0.3–0.6 mm, adaxial nectary oblong or ovate, 0.4–1.2 mm, nectaries distinct;

filaments distinct, hairy basally or on proximal 1/2;

anthers globose, 0.4–0.6 mm.

Pistillate flowers

abaxial nectary (0–)0.8–0.9 mm, adaxial nectary oblong, ovate, or narrowly oblong to almost filiform, 0.7–1.4 mm, shorter to longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0.4–1.2 mm;

ovary pyriform or obnapiform, densely short-silky, hairs cylindrical, beak abruptly tapering to styles;

ovules 13–15 per ovary;

styles connate 1/2 their lengths to almost distinct, 0.2–0.4 mm;

stigmas flat, abaxially non-papillate with rounded tip, slenderly cylindrical, or 2 plump lobes, 0.2–0.28–0.36 mm.

abaxial nectary absent, adaxial nectary oblong, 0.4–1.3 mm, shorter than stipe;

stipe 0.8–1.3 mm;

ovary pyriform, very densely long-silky, beak gradually tapering to or slightly bulged below styles;

ovules 6–14 per ovary;

styles 0.5–1 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly to slenderly cylindrical, 0.28–0.35–0.48 mm.

Capsules

3–5 mm.

5.6–7.2 mm.

Catkins

staminate 13–48 × 4–6.5(–8) mm, flowering branchlet 3–31(–50) mm; pistillate densely flowered, slender or stout, 18–56 × 4–10 mm, flowering branchlet 3–27(–40) mm;

floral bract tawny, 0.8–1.6 mm, apex rounded, entire, abaxially densely hairy, hairs straight.

staminate 14–31 × 8–18 mm, flowering branchlet 1.5–9 mm; pistillate loosely flowered, stout to globose, 10–58 × 7–25 mm, flowering branchlet 3.5–26 mm;

floral bract tawny, 1–1.6 mm, apex rounded, entire, abaxially sparsely hairy, almost glabrous, hairs wavy.

2n

= 38.

= 76, 95, 114.

Salix vestita

Salix athabascensis

Phenology Flowering mid Jun-late Jul. Flowering late May-late Jul.
Habitat Moist to dry open forests and rocky streamsides, in upper montane and subalpine zones, rarely alpine Fens, bogs, and treed bogs
Elevation 0-2400 m (0-7900 ft) 0-1800 m (0-5900 ft)
Distribution
from FNA
MT; OR; WA; AB; BC; MB; NL; NS; NU; ON; QC; Asia (China [Xinjiang], Mongolia, Russia, e, c Siberia)
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AB; BC; MB; NT; SK; YT
[BONAP county map]
Discussion

Salix vestita is an ancient amphiberingian species. Its distribution includes a series of isolated, disjunct populations in Central Siberia, the northern Rocky Mountains, the west coast of Hudson Bay, and the northeastern arctic and subarctic. Occurrence in Nunavut is on Akpatok Island in Ungava Bay and on the Belcher Islands in Hudson Bay. It may be extirpated in Washington.

The flowering and vegetative branchlets sometimes have relatively short internodes. In subsequent years, branches have the appearance of short shoots similar to those in Alnus. Short shoots do not appear on all branches or in all years. The formation of short shoots may be related to adverse growing conditions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The three polyploid chromosome numbers reported for Salix athabascensis, as well as the presence of leaves with ferruginous hairs, otherwise unknown in sect. Myrtilloides, are indicators of allopolyploidy.

Hybrids:

Salix athabascensis forms natural hybrids with S. pedicellaris. These hybrids combine the characteristics of the parents. The ovaries may be moderately densely villous or glabrous, but commonly have hairs in patches, or the stipes may be hairy and the ovaries glabrous; juvenile blades, and sometimes mature leaves, are hairy with white and ferruginous hairs. Some plants that resemble S. athabascensis have leaves glaucous adaxially, as in S. pedicellaris. The ovaries often appear to be infertile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 64. FNA vol. 7, p. 84.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Chamaetia > sect. Myrtilloides
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. leiolepis, S. vestita subsp. leiolepis, S. vestita var. psilophylla S. fallax, S. pedicellaris var. athabascensis
Name authority Pursh: Fl. Amer. Sept. 2: 610. (1813) Raup: Rhodora 32: 111, plate 202. (1930)
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