Salix uva-ursi |
Salix serissima |
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bearberry willow |
autumn willow |
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Habit | Plants 0.01–0.05 m, (dwarf), forming clonal compact mats by layering. | Shrubs, 1–5 m. Stems: branches usually flexible at base, sometimes brittle, yellow-brown, red-brown, or gray-brown, glabrous, slightly glossy or dull; branchlets yellow-brown or red-brown, glabrous, slightly or highly glossy. |
Stems | prostrate, short-trailing; branches red-brown, gray-brown, or yellow-brown, glabrous; branchlets yellow-green or yellow-brown, glabrous or puberulent. |
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Leaves | stipules (sometimes marcescent) absent, rudimentary, or foliaceous on early ones, foliaceous on late ones; petiole (shallowly to deeply grooved adaxially), 2–6.5 mm; largest medial blade (marcescent), amphistomatous or hypostomatous, ovate, broadly obovate, oblanceolate, or elliptic, 4–23 × 3.5–10 mm, 1.7–3.6 times as long as wide, base convex or cuneate, margins flat, serrulate or crenulate, apex convex, acuminate, acute, or retuse, abaxial surface glaucous, usually glabrous (rarely few hairs), adaxial slightly or highly glossy, usually glabrous (rarely a few hairs); proximal blade margins entire or serrate; juvenile blade glabrous, pilose, or puberulent abaxially. |
stipules absent or rudimentary; petiole shallowly to deeply grooved adaxially, 3–13 mm, with pairs of spherical glands distally or throughout, glabrous adaxially; largest medial blade hypostomatous or hemiamphistomatous, narrowly oblong, very narrowly elliptic, elliptic, lanceolate, or narrowly ovate, 43–110 × 9–33 mm, 2.4–6 times as long as wide, base convex or cuneate, margins flat, serrulate, apex acuminate, caudate, or acute, abaxial surface usually not glaucous, sometimes thinly so (appearing pale green), slightly glossy, glabrous, adaxial highly glossy, glabrous; proximal blade margins serrulate or entire; juvenile blade reddish or yellowish green, glabrous abaxially. |
Staminate flowers | abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.4–0.9 mm; filaments distinct, glabrous; stamens usually 1, rarely 2; anthers ellipsoid or shortly cylindrical, 0.4–0.7 mm. |
abaxial nectary 0.5–1.1 mm, adaxial nectary oblong or ovate, 0.4–1.1 mm, nectaries distinct or connate and cup-shaped; stamens 3–9; filaments distinct or basally connate, hairy on proximal 1/2 or basally; anthers ellipsoid or shortly cylindrical, 0.5–0.7 mm. |
Pistillate flowers | abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.5–0.8 mm, shorter to longer than stipe; stipe 0.3–1.6 mm; ovary ovoid or pyriform, glabrous, beak gradually tapering to styles; ovules 4–9 per ovary; styles 0.4–1 mm; stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.1–0.23–0.4 mm. |
adaxial nectary ovate, oblong, or flask-shaped, 0.3–1.1 mm, shorter than stipe; stipe 1.2–2.4 mm; ovary pyriform to obclavate, beak slightly bulged below or abruptly tapering to styles; ovules 12–16 per ovary; styles connate, 0.3–1 mm; stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.4–0.7 mm. |
Capsules | 3–5 mm. |
7–12 mm. |
Catkins | staminate 9–19 × 5–8 mm, flowering branchlet 0.5–9 mm; pistillate densely flowered, slender to subglobose, 11–47(–55 in fruit) × 6–10 mm, flowering branchlet 2–10 mm; floral bract brown, black, tawny, light rose, or bicolor, 1.1–1.8 mm, apex rounded or acute, entire, abaxially sparsely hairy, hairs straight or wavy. |
staminate (stout), 25–53 × 12–16 mm, flowering branchlet 5–14 mm; pistillate (fruiting in autumn, often persistent) moderately densely to loosely flowered, stout to globose, 17–42(–65 in fruit) × 11–22 mm, flowering branchlet 5–32(–65 in fruit) mm; floral bract (sometimes greenish tawny), 1.2–4 mm, apex acute, rounded, or truncate, glandular-toothed, abaxially moderately densely hairy, hairs straight or wavy. |
2n | = 38. |
= 76. |
Salix uva-ursi |
Salix serissima |
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Phenology | Flowering mid Jun-early Aug. | Flowering early Jun-early Jul. |
Habitat | Exposed, often dry or moist, calcareous, serpentine, dioritic, and granitic rocks, boulders, gravel, sand on beaches, outcrops, in snowbeds | Wet thickets, fens, brackish marshy strands, marly lakeshores, treed bogs, gravelly stream banks, lakeshores |
Elevation | 10-1200 m (0-3900 ft) | 10-3000 m (0-9800 ft) |
Distribution |
ME; NH; NY; VT; NL; NS; NU; QC; SPM; Greenland
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CO; CT; IL; IN; MA; MI; MN; MT; ND; NJ; NY; OH; PA; SD; VT; WI; WY; AB; BC; MB; NB; NL; NT; NU; QC; SK
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Discussion | Hybrids: Salix uva-ursi forms natural hybrids with S. herbacea. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Salix serissima is found in Nunavut only on Akimiski Island in James Bay. Flowering of Salix serissima is often described as serotinous (i.e., long after leaves emerge), but actually, they flower just as leaves emerge. Although they flower only a little later in spring than related species, they set fruit in late summer, and fruiting catkins often persist throughout winter. Their seeds remain dormant throughout the winter and germinate in the spring, thus enabling them to invade fens by completing their first annual growth before the sedges and grasses are tall enough to shade them out. This strategy has been reported to occur also in the related S. pentandra (A. K. Skvortsov 1999). North American Salix serissima is closely related to Eurasian S. pseudopentandra (Floderus) Floderus (A. K. Skvortsov 1999), which is known in China as S. pentandra var. intermedia Nakai and possibly also S. humaensis Y. L. Chou & R. C. Chou (Fang Z. F. et al. 1999). The relationship of S. serissima and S. pseudopentandra is similar to that of S. arbusculoides and S. boganidensis (G. W. Argus 1997). These two species, along with the amphiberingian S. vestita, are relictual members of former panboreal distributions. Hybrids: Hybrids between Salix lucida and S. serissima have been reported (M. L. Fernald 1950); no convincing specimens have been seen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 73. | FNA vol. 7, p. 45. |
Parent taxa | Salicaceae > Salix > subg. Chamaetia > sect. Myrtosalix | Salicaceae > Salix > subg. Salix > sect. Salicaster |
Sibling taxa | ||
Synonyms | S. ivigtutiana, S. myrsinites var. parvifolia | S. lucida var. serissima |
Name authority | Pursh: Fl. Amer. Sept. 2: 610. (1813) | (L. H. Bailey) Fernald: Rhodora 6: 6. (1903) |
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