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bearberry willow

bayberry willow, blue-leaf willow

Habit Plants 0.01–0.05 m, (dwarf), forming clonal compact mats by layering. Plants 0.3–5 m, (sometimes forming clones by stem fragmentation or layering).
Stems

prostrate, short-trailing;

branches red-brown, gray-brown, or yellow-brown, glabrous;

branchlets yellow-green or yellow-brown, glabrous or puberulent.

branches (sometimes highly brittle at base), red-brown or yellow-brown, not or weakly glaucous, (slightly or highly glossy), glabrous or villous;

branchlets red-brown or yellow-brown, glabrous or sparsely to very densely villous, (buds caprea-type, inner membranaceous bud-scale layer free and separating or not).

Leaves

stipules (sometimes marcescent) absent, rudimentary, or foliaceous on early ones, foliaceous on late ones;

petiole (shallowly to deeply grooved adaxially), 2–6.5 mm;

largest medial blade (marcescent), amphistomatous or hypostomatous, ovate, broadly obovate, oblanceolate, or elliptic, 4–23 × 3.5–10 mm, 1.7–3.6 times as long as wide, base convex or cuneate, margins flat, serrulate or crenulate, apex convex, acuminate, acute, or retuse, abaxial surface glaucous, usually glabrous (rarely few hairs), adaxial slightly or highly glossy, usually glabrous (rarely a few hairs);

proximal blade margins entire or serrate;

juvenile blade glabrous, pilose, or puberulent abaxially.

stipules rudimentary or foliaceous on early ones, foliaceous on late ones, apex acute or acuminate;

petiole shallowly grooved, or convex to flat adaxially, 3.5–7.3–13 mm, (sometimes with 2 spherical glands distally), villous, tomentose, pilose, or pubescent adaxially;

largest medial blade narrowly oblong, narrowly elliptic, elliptic, or oblanceolate, 35–61.3–110 × 11–16–46 mm, 2–2.7–5.2 times as long as wide, base convex, rounded, subcordate, or cuneate, margins flat or slightly revolute, (thickened and raised), crenulate or serrulate, apex acuminate, acute, or convex, abaxial surface usually very thickly glaucous, glabrous or pilose, midribs pubescent to tomentose, hairs (white, often also ferruginous), curved, wavy, or straight, adaxial slightly glossy, glabrous or pilose, midribs sparsely pubescent (hairs white, sometimes also ferruginous);

proximal blade margins entire or serrulate;

juvenile blade translucent, reddish or yellowish green, glabrous or sparsely pubescent abaxially, midribs often densely hairy, hairs white, sometimes also ferruginous.

Staminate flowers

abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.4–0.9 mm;

filaments distinct, glabrous;

stamens usually 1, rarely 2;

anthers ellipsoid or shortly cylindrical, 0.4–0.7 mm.

adaxial nectary narrowly oblong, oblong, square, or ovate, 0.44–0.48–1.4 mm;

filaments distinct, glabrous;

anthers yellow, (ellipsoid or globose), 0.52–0.69–0.76 mm.

Pistillate flowers

abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.5–0.8 mm, shorter to longer than stipe;

stipe 0.3–1.6 mm;

ovary ovoid or pyriform, glabrous, beak gradually tapering to styles;

ovules 4–9 per ovary;

styles 0.4–1 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.1–0.23–0.4 mm.

adaxial nectary oblong, narrowly oblong, square, or flask-shaped, 0.56–0.8–1.4 mm, shorter than stipe;

stipe 0.96–1.7–3.4 mm;

ovary pyriform, glabrous, beak slightly bulged below styles;

ovules 12–14 per ovary;

styles (sometimes distinct), 0.3–0.8–1.3 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or slenderly cylindrical, 0.24–0.43–0.56 mm.

Capsules

3–5 mm.

5–7–11 mm.

Catkins

staminate 9–19 × 5–8 mm, flowering branchlet 0.5–9 mm; pistillate densely flowered, slender to subglobose, 11–47(–55 in fruit) × 6–10 mm, flowering branchlet 2–10 mm;

floral bract brown, black, tawny, light rose, or bicolor, 1.1–1.8 mm, apex rounded or acute, entire, abaxially sparsely hairy, hairs straight or wavy.

staminate flowering before leaves emerge, pistillate as leaves emerge; staminate stout or slender, 23.5–35.6–51 × 9–12.7–22 mm, flowering branchlet 1–3.9–10 mm; pistillate loosely flowered, stout or slender, 19–42–62(–85 in fruit) × 8–13–18 mm, flowering branchlet 1.5–5.9–13 mm;

floral bract brown or bicolor, 1.2–1.8–3 mm, apex rounded or acute, sometimes toothed, abaxially hairy, hairs straight or wavy.

2n

= 38.

Salix uva-ursi

Salix myricoides

Phenology Flowering mid Jun-early Aug. Flowering early Apr-early Jul.
Habitat Exposed, often dry or moist, calcareous, serpentine, dioritic, and granitic rocks, boulders, gravel, sand on beaches, outcrops, in snowbeds Stream and lake shores, gravel bars, subalpine conifer forests, alkaline fens, sea cliffs, dry limestone talus, swamps, tidal meadows, sand dunes
Elevation 10-1200 m (0-3900 ft) 0-1100 m (0-3600 ft)
Distribution
from FNA
ME; NH; NY; VT; NL; NS; NU; QC; SPM; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
IL; IN; ME; MI; OH; PA; WI; NB; NF; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Hybrids:

Salix uva-ursi forms natural hybrids with S. herbacea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants with densely villous branchlets and branches have been named var. albovestita. Branchlet indumentum varies widely in the species and seems to be continuous, with both villous and glabrous variants sometimes occurring in the same area. This characteristic is more common in populations on the shores of the Great Lakes and on the western coast of James Bay, but even these populations are variable.

Reports of Salix myricoides from Akimiski Island, Nunavut, were based on misidentified S. planifolia. See 68. S. eriocephala for differences.

Hybrids:

Salix myricoides forms natural hybrids with S. bebbiana, S. discolor, and S. glauca var. cordifolia. Hybrids with S. eriocephala have been reported (M. L. Fernald 1950) but no convincing specimens have been seen. Inasmuch as S. eriocephala and S. myricoides are very similar, hybrids between the two would be very difficult to identify.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 73. FNA vol. 7, p. 119.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Myrtosalix Salicaceae > Salix > subg. Vetrix > sect. Hastatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. ivigtutiana, S. myrsinites var. parvifolia S. glaucophylla var. albovestita, S. glaucophylloides, S. glaucophylloides var. albovestita, S. glaucophylloides var. glaucophylla, S. myricoides var. albovestita
Name authority Pursh: Fl. Amer. Sept. 2: 610. (1813) Muhlenberg: Ges. Naturf. Freunde Berlin Neue Schriften 4: 235, plate 6, fig. 2. (1803)
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