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bearberry willow

Del Norte willow

Habit Plants 0.01–0.05 m, (dwarf), forming clonal compact mats by layering. Shrubs, 1–2 m, (sometimes forming clones by stem fragmentation).
Stems

prostrate, short-trailing;

branches red-brown, gray-brown, or yellow-brown, glabrous;

branchlets yellow-green or yellow-brown, glabrous or puberulent.

branches (highly brittle at base), red-brown, not glaucous, tomentose or velvety to glabrescent;

branchlets red-brown or yellow-brown, densely velvety, (buds caprea-type).

Leaves

stipules (sometimes marcescent) absent, rudimentary, or foliaceous on early ones, foliaceous on late ones;

petiole (shallowly to deeply grooved adaxially), 2–6.5 mm;

largest medial blade (marcescent), amphistomatous or hypostomatous, ovate, broadly obovate, oblanceolate, or elliptic, 4–23 × 3.5–10 mm, 1.7–3.6 times as long as wide, base convex or cuneate, margins flat, serrulate or crenulate, apex convex, acuminate, acute, or retuse, abaxial surface glaucous, usually glabrous (rarely few hairs), adaxial slightly or highly glossy, usually glabrous (rarely a few hairs);

proximal blade margins entire or serrate;

juvenile blade glabrous, pilose, or puberulent abaxially.

stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 6–16 mm, velvety or tomentose adaxially;

largest medial blade elliptic or obovate, 53–102 × 29–54 mm, 1.3–2.8 times as long as wide, base cuneate or convex, margins slightly revolute, entire or sinuate, apex convex, rounded, or acute, abaxial surface glaucous (sometimes obscured by hairs), densely to sparsely velvety, tomentose, villous, or short-silky, hairs erect or spreading, wavy, adaxial dull, sparsely tomentose or short-silky;

proximal blade margins entire;

juvenile blade green, very densely velvety or long-silky abaxially, hairs white or gray.

Staminate flowers

abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.4–0.9 mm;

filaments distinct, glabrous;

stamens usually 1, rarely 2;

anthers ellipsoid or shortly cylindrical, 0.4–0.7 mm.

adaxial nectary narrowly oblong to oblong, 0.5–1 mm;

filaments distinct or slightly connate, glabrous;

anthers purple turning yellow, ellipsoid, 0.6–0.7 mm.

Pistillate flowers

abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.5–0.8 mm, shorter to longer than stipe;

stipe 0.3–1.6 mm;

ovary ovoid or pyriform, glabrous, beak gradually tapering to styles;

ovules 4–9 per ovary;

styles 0.4–1 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.1–0.23–0.4 mm.

adaxial nectary narrowly oblong to oblong, 0.4–0.9 mm, longer than stipe;

stipe 0–0.3 mm;

ovary obturbinate, short-silky or densely pubescent, beak gradually tapering to styles;

ovules 14–18 per ovary;

styles 0.6–1.2 mm;

stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.2–0.32–0.4 mm.

Capsules

3–5 mm.

4 mm.

Catkins

staminate 9–19 × 5–8 mm, flowering branchlet 0.5–9 mm; pistillate densely flowered, slender to subglobose, 11–47(–55 in fruit) × 6–10 mm, flowering branchlet 2–10 mm;

floral bract brown, black, tawny, light rose, or bicolor, 1.1–1.8 mm, apex rounded or acute, entire, abaxially sparsely hairy, hairs straight or wavy.

flowering before leaves emerge; staminate stout, 25–30 × 6–13 mm, flowering branchlet 0–5 mm; pistillate moderately densely flowered, slender to stout, 17–53 × 6–8 mm, flowering branchlet 1–3 mm;

floral bract brown, 1.2–2.2 mm, apex rounded, abaxially hairy, hairs straight.

2n

= 38.

Salix uva-ursi

Salix delnortensis

Phenology Flowering mid Jun-early Aug. Flowering late Mar-early May.
Habitat Exposed, often dry or moist, calcareous, serpentine, dioritic, and granitic rocks, boulders, gravel, sand on beaches, outcrops, in snowbeds Streamshores, gravel to boulder substrates, serpentine soils
Elevation 10-1200 m (0-3900 ft) 90-500 m (300-1600 ft)
Distribution
from FNA
ME; NH; NY; VT; NL; NS; NU; QC; SPM; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Hybrids:

Salix uva-ursi forms natural hybrids with S. herbacea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The origin of the serpentine endemics Salix breweri and S. delnortensis is still unresolved (G. W. Argus 1997; R. D. Dorn 2000). The possibility that they are of hybrid origin, perhaps involving S. lasiolepis and S. sitchensis (Dorn 1976), cannot be resolved by field and herbarium studies alone; cytological and experimental methods need to be employed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 73. FNA vol. 7, p. 161.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Myrtosalix Salicaceae > Salix > subg. Vetrix > sect. Sitchenses
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. ivigtutiana, S. myrsinites var. parvifolia S. breweri var. delnortensis
Name authority Pursh: Fl. Amer. Sept. 2: 610. (1813) C. K. Schneider: J. Arnold Arbor. 1: 96. (1919)
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