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creeping willow, sprouting leaf willow

dark-leaf willow, myrsine-leaf willow

Habit Plants 0.02–0.1 m, forming clones by layering or rhizomes.
Stems

trailing or erect;

branches red-brown, (sometimes weakly glaucous), glabrous;

branchlets yellow-brown or greenish brown, glabrous.

branches dark red-brown or gray-brown, not glaucous, glabrous or hairy, (peeled wood smooth or striate with relatively few, short striae);

branchlets red-brown, moderately to very densely pubescent or velvety.

Leaves

stipules (sometimes marcescent) absent or rudimentary on early ones, rudimentary or foliaceous on late ones;

petiole (deeply to shallowly grooved adaxially), 3–9–20 mm, (ciliate, glabrous adaxially);

largest medial blade amphistomatous or hemiamphistomatous, (sometimes with 2 pairs of secondary veins arising at or close to base, arcing toward apex), elliptic, broadly elliptic, or subcircular, 16–42 × 12–30(–38) mm, 1–2 times as long as wide, base convex, cuneate, or rounded, margins flat or slightly revolute, entire or serrulate, ciliate (hairs wavy), apex convex, acuminate, rounded, or retuse, abaxial surface glabrous, adaxial highly glossy, glabrous;

proximal blade margins entire or irregularly serrulate;

juvenile blade pilose to glabrescent abaxially.

stipules usually foliaceous, sometimes minute rudiments on early ones, foliaceous on late ones, (ca. 4 mm), apex acute;

petiole convex to flat, or shallowly grooved adaxially, 3.5–12(–15) mm, villous to puberulent adaxially;

largest medial blade (sometimes hemiamphistomatous), broadly obovate, elliptic, broadly elliptic, or subcircular, 24–52(–100) × 12–45 mm, base concave, rounded, subcordate, cordate, or cuneate, margins sometimes slightly revolute, serrulate, or crenulate to subentire, apex abruptly acuminate or acute, abaxial surface glaucous (tip often not glaucous), sparsely to moderately densely puberulent, or silky to glabrescent, hairs appressed or spreading, straight or wavy, adaxial slightly glossy, glabrescent or sparsely to moderately densely puberulent, pubescent, or short-silky (especially midrib);

proximal blade margins serrulate, crenulate, or entire;

juvenile blade sometimes reddish, long-silky, villous, tomentose (at least on midrib), or glabrous abaxially, hairs usually white, rarely somewhat ferruginous.

Staminate flowers

abaxial nectary (0–)0.2–0.7 mm, adaxial nectary oblong, narrowly oblong, or ovate, 0.6–1.3 mm, nectaries distinct;

filaments distinct (glabrous);

anthers ellipsoid, 0.5–0.6 mm.

adaxial nectary 0.5–0.7 mm;

filaments distinct;

anthers purple turning yellow, (ellipsoid or shortly cylindrical), 0.5–0.8 mm.

Pistillate flowers

abaxial nectary absent, adaxial nectary oblong, 0.5–1.4 mm, longer than stipe;

stipe 0.2–0.8 mm;

ovary pyriform, sometimes glaucous, glabrous, beak gradually tapering to styles;

ovules 12–13 per ovary;

styles connate to distinct 1/2 their lengths, (0.6–)0.8–2 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or slenderly cylindrical, 0.32–0.5–0.88 mm.

adaxial nectary ovate, square, or flask-shaped, 0.4–0.6(–1) mm;

ovary pyriform, pubescent throughout or in patches or streaks, or glabrous (hairs refractive), beak gradually tapering to styles;

ovules 12–14 per ovary;

styles 0.6–1.5 mm.

Capsules

4–10 mm.

6–10 mm.

Catkins

staminate 13–31 × 7–11 mm, flowering branchlet 1–15 mm;

pistillate moderately densely to loosely flowered, stout to globose, 15–54(–90 in fruit) × 6–15 mm, flowering branchlet 2–42 mm;

floral bract brown, 1.6–2 mm, apex rounded, entire, abaxially sparsely hairy throughout or distally, hairs straight or wavy.

flowering as leaves emerge;

staminate (densely flowered), stout or subglobose, 17–35 mm, flowering branchlet 1–3 mm;

pistillate densely flowered, stout, 9–11(–30)(–80 in fruit) mm, flowering branchlet 2–5.5 mm;

floral bract pale brown, 1–1.9(–2.8) mm, apex acute, convex, or rounded, abaxially sparsely hairy, hairs straight or wavy.

2n

= 114.

Salix stolonifera

Salix myrsinifolia

Phenology Flowering early Jun-early Jul. Flowering mid Mar-mid Jun.
Habitat Arctic, subarctic, and alpine, wet sedge meadows, hummocky tundra, raised center polygons, Dryas-willow-sedge tundra, Dryas mats on dry ridge tops Roadsides, waste places
Elevation 0-1000 m (0-3300 ft) ca. 100 m (ca. 300 ft)
Distribution
from FNA
AK; AB; BC
[BONAP county map]
from FNA
ON; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Hybrids:

Salix stolonifera forms natural hybrids with S. arctica and S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix myrsinifolia may be naturalized in the vicinity of Ottawa, Ontario, but that needs confirmation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 75. FNA vol. 7, p. 125.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Ovalifoliae Salicaceae > Salix > subg. Vetrix > sect. Nigricantes
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. nigricans
Name authority Coville: Proc. Wash. Acad. Sci. 3: 333, plate 41, fig. 1. (1901) Salisbury: Prodr. Stirp. Chap. Allerton, 394. (1796)
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