Salix sitchensis |
Salix turnorii |
|
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Coulter willow, Sitka willow |
Turnor's willow |
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Habit | Shrubs or trees, 1–8 m, (sometimes forming clones by stem fragmentation). | Shrubs, 1–2.5 m, (forming clones by layering). |
Stems | branches (sometimes highly brittle at base), yellow-brown or red-brown, not glaucous, glabrous or pilose; branchlets yellow-brown, gray-brown, or red-brown, densely short-silky, velvety, or villous, (buds caprea-type or intermediate). |
branches yellow-brown or yellow-gray, not or weakly glaucous, (with sparkling wax crystals, dull or slightly glossy), pilose or villous; branchlets gray-brown or red-brown, pubescent, villous, or velvety. |
Leaves | stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute; petiole convex to flat, or shallowly grooved adaxially, 3–13(–16) mm, tomentose or velvety adaxially; largest medial blade elliptic, narrowly oblanceolate, oblanceolate, or obovate, 31–70–120 × 17–48 mm, 2.1–3.1–4 times as long as wide, base cuneate or convex, margins slightly revolute or flat, strongly revolute proximally, entire, irregularly serrate, or sinuate, (glands submarginal or epilaminal), apex acuminate or convex, abaxial surface not evidently glaucous, (obscured by hairs), densely short-silky, woolly, or silky-woolly, hairs straight, wavy, or curved, adaxial slightly glossy (sometimes dull and glaucous), pilose or moderately densely short-silky; proximal blade margins entire or shallowly serrulate; juvenile blade green, densely long-silky or woolly abaxially, (sparsely silky-tomentose adaxially), hairs white. |
stipules foliaceous, apex acute or convex; petiole shallowly grooved adaxially, 4–13 mm, villous or pubescent adaxially; largest medial blade (amphistomatous), narrowly oblong, narrowly elliptic, elliptic, oblanceolate, or lanceolate, 26–47 × 7.5–15 mm, 2.8–4.1 times as long as wide, base convex, rounded, or subcordate, margins slightly revolute or flat, serrate or serrulate, apex acuminate to acute, abaxial surface glaucous, glabrous, pilose, villous, or long-silky, hairs straight or wavy, adaxial dull, sparsely or moderately densely pilose or long-silky, especially on midrib; proximal blade margins entire or serrulate; juvenile blade reddish or yellowish green, sparsely to moderately densely long-silky or pubescent abaxially, hairs white. |
Staminate flowers | adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.4–1.3 mm; stamens 1; filaments distinct, glabrous; anthers purple turning yellow, shortly cylindrical, 0.5–0.7 mm. |
adaxial nectary narrowly oblong or ovate, 0.7–1.1 mm; filaments distinct or connate, hairy basally; anthers yellow, 0.6–0.8 mm. |
Pistillate flowers | adaxial nectary square, ovate, or flask-shaped, 0.5–0.9 mm, shorter to longer than stipe; stipe 0.4–1.4 mm; ovary ovoid to pyriform, long- or short-silky or villous, beak sometimes slightly bulged below styles; ovules 14–20 per ovary; styles 0.4–0.8 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, 0.16–0.28–0.4 mm. |
adaxial nectary oblong, flask-shaped, or narrowly ovate, 0.4–1 mm, shorter than stipe; stipe 2–4 mm; ovary pyriform, glabrous, beaks slightly bulged below styles; ovules 14–18 per ovary; styles 0.3–0.5 mm; stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.16–0.23–0.28 mm. |
Capsules | 3.5–5.6 mm. |
2.5–5 mm. |
Catkins | flowering just before or as leaves emerge; staminate slender or stout, (17–)22–54 × 8–15 mm, flowering branchlet 1–9 mm; pistillate moderately densely flowered, slender to stout, 25–73(–115 in fruit) × 5–15 mm, flowering branchlet 1–20 mm; floral bract tawny to dark brown, 1.4–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight or wavy. |
flowering just before or as leaves emerge; staminate stout, 16–30 × 8–10 mm, flowering branchlet 1.5–4 mm; pistillate loosely flowered, stout, 18–22 × 9–11 mm, flowering branchlet 3–4 mm; floral bract brown or tawny, 1.2–1.6 mm, apex acute or rounded, abaxially hairy throughout or proximally, hairs straight or wavy. |
2n | = 38. |
|
Salix sitchensis |
Salix turnorii |
|
Phenology | Flowering early Apr-mid Jun (Mar in California). | No flowering time data are available (probably May or Jun). |
Habitat | Tidal swamps and marshes, coastal fog belts and headlands, sand dunes, springs, gravelly streambeds and deltas, glacial moraines, avalanche tracks, dry canyons, clearings and edges of forests, shade tolerant | Active sand dunes |
Elevation | 0-1800 m (0-5900 ft) | 200-300 m (700-1000 ft) |
Distribution |
AK; CA; ID; MT; OR; WA; AB; BC
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SK |
Discussion | Ovary hairiness in some Salix sitchensis populations varies from uniformly hairy to glabrescent, with intermediates with patchy or streaky hairiness. All three variations can occur together and do not seem to indicate hybridization. Both Salix sitchensis and S. scouleriana have similar variants with leaves having very densely curly hairs on abaxial surfaces [S. sitchensis forma coulteri (Andersson) Jepson and S. scouleriana forma poikila (C. K. Schneider) C. K. Schneider]. Plants resembling S. drummondiana but with similar indumentum probably are hybrids with S. alaxensis (see 84. S. drummondiana). The coulteri taxon resembles S. delnortensis in having stipules with adaxial surfaces glabrous and very sparsely glandular toward the base, densely hairy abaxially, and with gland-dotted margins; its branchlets have wavy to crinkly hairs. The possible hybrid origin of S. delnortensis needs study (R. D. Dorn 2000). Hybrids: Salix sitchensis forms natural hybrids with S. alaxensis var. longistylis and S. melanopsis. Hybridization with S. geyeriana reported by J. K. Henry (1915) is not based on convincing specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Salix turnorii is known from the Lake Athabasca sand dunes in northwestern Saskatchewan. Salix famelica in the Great Sand Hills, southern Saskatchewan, is very similar morphologically and may have been the source of populations ancestral to S. turnorii that moved into northern Saskatchewan during the warm Holocene Hypsithermal Period (ca. 9000–6000 yrs. B.P.). Hybrids: Salix turnorii forms natural hybrids with S. brachycarpa var. psammophila. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 159. | FNA vol. 7, p. 122. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Sitchenses | Salicaceae > Salix > subg. Vetrix > sect. Cordatae |
Sibling taxa | ||
Synonyms | S. coulteri, S. sitchensis var. parvifolia | S. lutea var. turnorii |
Name authority | Sanson ex Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 162. (1832) | Raup: J. Arnold Arbor. 17: 234, plate 193. (1936) |
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