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Coulter willow, Sitka willow

Richardson's willow

Habit Shrubs or trees, 1–8 m, (sometimes forming clones by stem fragmentation). Plants 0.3–6.5 m.
Stems

branches (sometimes highly brittle at base), yellow-brown or red-brown, not glaucous, glabrous or pilose;

branchlets yellow-brown, gray-brown, or red-brown, densely short-silky, velvety, or villous, (buds caprea-type or intermediate).

branches red-brown, yellow-brown, or violet, not to strongly glaucous (slightly or highly glossy), villous to glabrescent with persistent patches of hairs;

branchlets red-brown or yellow-brown, pilose to densely villous, (buds caprea-type, scale inner membrane free, separating from outer layer).

Leaves

stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 3–13(–16) mm, tomentose or velvety adaxially;

largest medial blade elliptic, narrowly oblanceolate, oblanceolate, or obovate, 31–70–120 × 17–48 mm, 2.1–3.1–4 times as long as wide, base cuneate or convex, margins slightly revolute or flat, strongly revolute proximally, entire, irregularly serrate, or sinuate, (glands submarginal or epilaminal), apex acuminate or convex, abaxial surface not evidently glaucous, (obscured by hairs), densely short-silky, woolly, or silky-woolly, hairs straight, wavy, or curved, adaxial slightly glossy (sometimes dull and glaucous), pilose or moderately densely short-silky;

proximal blade margins entire or shallowly serrulate;

juvenile blade green, densely long-silky or woolly abaxially, (sparsely silky-tomentose adaxially), hairs white.

stipules (sometimes marcescent), foliaceous, (3–12–35 mm, 1.7–3.6–6.6 times as long as wide, lanceolate or narrowly elliptic, sometimes ovate or oval, base broad, abruptly tapering), apex acuminate or acute;

petiole convex to flat or shallowly grooved adaxially, 2–7.3–27 mm, villous to glabrescent adaxially;

largest medial blade broadly to narrowly elliptic, or obovate, 23–45–100 × 10–22–55 mm, 1.2–2.2–4.2 times as long as wide, base cuneate, convex, or concave, margins slightly revolute or flat, entire or serrulate, apex acute, acuminate, or convex, abaxial surface glaucous, glabrous or pilose, hairs (white, sometimes also ferruginous), curved, adaxial slightly glossy, moderately densely villous or pilose to glabrescent;

proximal blade margins usually entire, sometimes serrulate;

juvenile blade yellowish green, abaxially densely villous or pilose, sometimes glabrous, hairs white, sometimes also ferruginous.

Catkins

flowering just before or as leaves emerge;

staminate slender or stout, (17–)22–54 × 8–15 mm, flowering branchlet 1–9 mm;

pistillate moderately densely flowered, slender to stout, 25–73(–115 in fruit) × 5–15 mm, flowering branchlet 1–20 mm;

floral bract tawny to dark brown, 1.4–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight or wavy.

flowering before leaves emerge;

staminate stout or subglobose, 22–53 × 16–24 mm, flowering branchlet 0 mm;

pistillate densely flowered, slender or stout, 25–69(–85 in fruit) × 13–20 mm, flowering branchlet 0–2 mm;

floral bract brown or black, 2.4–3.2 mm, apex acute, rounded, or retuse, abaxially moderately densely hairy, hairs straight.

Staminate flowers

adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.4–1.3 mm;

stamens 1;

filaments distinct, glabrous;

anthers purple turning yellow, shortly cylindrical, 0.5–0.7 mm.

adaxial nectary narrowly oblong to oblong, 0.7–1.8 mm;

filaments distinct to connate ca. 1/2 their lengths, glabrous;

anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.5–0.8 mm.

Pistillate flowers

adaxial nectary square, ovate, or flask-shaped, 0.5–0.9 mm, shorter to longer than stipe;

stipe 0.4–1.4 mm;

ovary ovoid to pyriform, long- or short-silky or villous, beak sometimes slightly bulged below styles;

ovules 14–20 per ovary;

styles 0.4–0.8 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, 0.16–0.28–0.4 mm.

adaxial nectary narrowly oblong, oblong, or flask-shaped, 0.6–1.4 mm (2+-lobed), longer than stipe;

stipe 0.4–1 mm;

ovary pyriform, beak gradually tapering to styles;

ovules 22–37 per ovary;

styles 1–2.8 mm;

stigmas slenderly cylindrical, 0.32–0.63–0.86 mm.

Capsules

3.5–5.6 mm.

4.5–6.8 mm.

2n

= 38.

= 38.

Salix sitchensis

Salix richardsonii

Phenology Flowering early Apr-mid Jun (Mar in California). Flowering late May-early Jul.
Habitat Tidal swamps and marshes, coastal fog belts and headlands, sand dunes, springs, gravelly streambeds and deltas, glacial moraines, avalanche tracks, dry canyons, clearings and edges of forests, shade tolerant Arctic, subarctic, subalpine and boreal, stream terraces and floodplains, open forests, wet sedge meadows, fens, drumlin fields, gravel ridges, bogs, cliff ledges, snowbeds, silt, sand and gravel, calcareous or igneous substrates
Elevation 0-1800 m [0-5900 ft] 10-1800 m [30-5900 ft]
Distribution
from FNA
AK; CA; ID; MT; OR; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC; MB; NT; NU; YT; e Asia (Chukotka, Kamchatka, Russia [Anadyr], e Siberia)
[BONAP county map]
Discussion

Ovary hairiness in some Salix sitchensis populations varies from uniformly hairy to glabrescent, with intermediates with patchy or streaky hairiness. All three variations can occur together and do not seem to indicate hybridization.

Both Salix sitchensis and S. scouleriana have similar variants with leaves having very densely curly hairs on abaxial surfaces [S. sitchensis forma coulteri (Andersson) Jepson and S. scouleriana forma poikila (C. K. Schneider) C. K. Schneider]. Plants resembling S. drummondiana but with similar indumentum probably are hybrids with S. alaxensis (see 84. S. drummondiana). The coulteri taxon resembles S. delnortensis in having stipules with adaxial surfaces glabrous and very sparsely glandular toward the base, densely hairy abaxially, and with gland-dotted margins; its branchlets have wavy to crinkly hairs. The possible hybrid origin of S. delnortensis needs study (R. D. Dorn 2000).

Hybrids:

Salix sitchensis forms natural hybrids with S. alaxensis var. longistylis and S. melanopsis. Hybridization with S. geyeriana reported by J. K. Henry (1915) is not based on convincing specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybrids:

Salix richardsonii forms natural hybrids with S. barclayi and S. calcicola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 159. Treatment author: George W. Argus. FNA vol. 7, p. 144. Treatment author: George W. Argus.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Sitchenses Salicaceae > Salix > subg. Vetrix > sect. Lanatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. coulteri, S. sitchensis var. parvifolia S. lanata subsp. richardsonii, S. richardsonii var. mckeandii
Name authority Sanson ex Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 162. (1832) Hooker: Fl. Bor.-Amer. 2: 147, plate 182. (1838)
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