Salix sitchensis |
Salix irrorata |
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Coulter willow, Sitka willow |
blue-stem willow, dewy-stem willow |
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Habit | Shrubs or trees, 1–8 m, (sometimes forming clones by stem fragmentation). | Shrubs, 2–7 m, (multistemmed, sometimes forming clones by stem fragmentation). |
Stems | branches (sometimes highly brittle at base), yellow-brown or red-brown, not glaucous, glabrous or pilose; branchlets yellow-brown, gray-brown, or red-brown, densely short-silky, velvety, or villous, (buds caprea-type or intermediate). |
branches (highly brittle or not at base), red-brown to violet, usually glaucous, glabrous; branchlets yellow-brown, (strongly glaucous or not), glabrous, sparsely velvety, or tomentose. |
Leaves | stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute; petiole convex to flat, or shallowly grooved adaxially, 3–13(–16) mm, tomentose or velvety adaxially; largest medial blade elliptic, narrowly oblanceolate, oblanceolate, or obovate, 31–70–120 × 17–48 mm, 2.1–3.1–4 times as long as wide, base cuneate or convex, margins slightly revolute or flat, strongly revolute proximally, entire, irregularly serrate, or sinuate, (glands submarginal or epilaminal), apex acuminate or convex, abaxial surface not evidently glaucous, (obscured by hairs), densely short-silky, woolly, or silky-woolly, hairs straight, wavy, or curved, adaxial slightly glossy (sometimes dull and glaucous), pilose or moderately densely short-silky; proximal blade margins entire or shallowly serrulate; juvenile blade green, densely long-silky or woolly abaxially, (sparsely silky-tomentose adaxially), hairs white. |
stipules absent or rudimentary on early ones, foliaceous or rudimentary on late ones, apex acute; petiole convex to flat adaxially, 4–14 mm, velvety adaxially; largest medial blade lorate, narrowly oblong, narrowly elliptic, or narrowly oblanceolate, 47–115 × 8–22 mm, 3.5–7.7 times as long as wide, base cuneate or convex, margins flat to slightly revolute, entire and gland-dotted, serrulate or crenate, apex acuminate, acute, or convex, abaxial surface glaucous, glabrous, sparsely tomentose, or short-silky, hairs wavy, adaxial slightly to highly glossy, glabrous or pilose; proximal blade margins entire or serrulate; juvenile blade yellowish green or reddish, glabrous or sparsely villous abaxially, hairs white. |
Staminate flowers | adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.4–1.3 mm; stamens 1; filaments distinct, glabrous; anthers purple turning yellow, shortly cylindrical, 0.5–0.7 mm. |
adaxial nectary narrowly oblong to oblong, 0.3–0.8 mm; filaments connate less than to more than 1/2 their lengths; anthers yellow or purple turning yellow, ellipsoid or shortly cylindrical, 0.4–0.7 mm. |
Pistillate flowers | adaxial nectary square, ovate, or flask-shaped, 0.5–0.9 mm, shorter to longer than stipe; stipe 0.4–1.4 mm; ovary ovoid to pyriform, long- or short-silky or villous, beak sometimes slightly bulged below styles; ovules 14–20 per ovary; styles 0.4–0.8 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, 0.16–0.28–0.4 mm. |
adaxial nectary narrowly oblong to oblong, 0.3–0.7 mm; stipe 0.4–1 mm; ovary pyriform, beak slightly bulged below styles; ovules 9–12 per ovary; styles 0.3–0.9 mm; stigmas flat, abaxially non-papillate with rounded tip, 0.15–0.28–0.6 mm. |
Capsules | 3.5–5.6 mm. |
3.5–4 mm. |
Catkins | flowering just before or as leaves emerge; staminate slender or stout, (17–)22–54 × 8–15 mm, flowering branchlet 1–9 mm; pistillate moderately densely flowered, slender to stout, 25–73(–115 in fruit) × 5–15 mm, flowering branchlet 1–20 mm; floral bract tawny to dark brown, 1.4–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight or wavy. |
flowering before or just before leaves emerge; staminate stout or subglobose, 15–34 × 8–22 mm, flowering branchlet 0(–2) mm; pistillate densely flowered, stout or slender, 14–43 × 7–12 mm, flowering branchlet 0–4 mm; floral bract brown or black, 1.3–2.5 mm, apex rounded, abaxially hairy, hairs straight or wavy. |
2n | = 38. |
= 38. |
Salix sitchensis |
Salix irrorata |
|
Phenology | Flowering early Apr-mid Jun (Mar in California). | Flowering mid Mar-mid May. |
Habitat | Tidal swamps and marshes, coastal fog belts and headlands, sand dunes, springs, gravelly streambeds and deltas, glacial moraines, avalanche tracks, dry canyons, clearings and edges of forests, shade tolerant | Streams, wet meadows |
Elevation | 0-1800 m (0-5900 ft) | 1400-3000 m (4600-9800 ft) |
Distribution |
AK; CA; ID; MT; OR; WA; AB; BC
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AZ; CO; NM; WY; Mexico (Baja California, Chihuahua, Coahuila, Durango, Sonora)
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Discussion | Ovary hairiness in some Salix sitchensis populations varies from uniformly hairy to glabrescent, with intermediates with patchy or streaky hairiness. All three variations can occur together and do not seem to indicate hybridization. Both Salix sitchensis and S. scouleriana have similar variants with leaves having very densely curly hairs on abaxial surfaces [S. sitchensis forma coulteri (Andersson) Jepson and S. scouleriana forma poikila (C. K. Schneider) C. K. Schneider]. Plants resembling S. drummondiana but with similar indumentum probably are hybrids with S. alaxensis (see 84. S. drummondiana). The coulteri taxon resembles S. delnortensis in having stipules with adaxial surfaces glabrous and very sparsely glandular toward the base, densely hairy abaxially, and with gland-dotted margins; its branchlets have wavy to crinkly hairs. The possible hybrid origin of S. delnortensis needs study (R. D. Dorn 2000). Hybrids: Salix sitchensis forms natural hybrids with S. alaxensis var. longistylis and S. melanopsis. Hybridization with S. geyeriana reported by J. K. Henry (1915) is not based on convincing specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Salix irrorata is very closely related to S. lasiolepis. The two are here maintained as separate species primarily because S. irrorata is a diploid and S. lasiolepis a tetraploid, and also because of their largely allopatric ranges (G. W. Argus 2007). Studies of sawflies (Euura Newman) by P. W. Price (pers. comm.) show that the same species of Euura can successfully reproduce on either willow. The only morphological character that separates the two is that in S. irrorata branchlets and branches are very strongly glaucous, whereas in S. lasiolepis they are not glaucous. Some plants have weakly glaucous stems (wax not visible except by polishing or only as sparkling crystals); this may be infraspecific variability or evidence of hybridization. Hybrids: Salix irrorata forms natural hybrids with S. drummondiana, S. geyeriana, and S. lasiolepis var. lasiolepis. Salix irrorata × S. lasiolepis var. lasiolepis: This putative hybrid occurs in Arizona and New Mexico. It is characterized mainly by weakly glaucous branches. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 159. | FNA vol. 7, p. 155. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Sitchenses | Salicaceae > Salix > subg. Vetrix > sect. Mexicanae |
Sibling taxa | ||
Synonyms | S. coulteri, S. sitchensis var. parvifolia | |
Name authority | Sanson ex Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 162. (1832) | Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 117. (1858) |
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