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Coulter willow, Sitka willow

Florida willow

Habit Shrubs or trees, 1–8 m, (sometimes forming clones by stem fragmentation).
Stems

branches (sometimes highly brittle at base), yellow-brown or red-brown, not glaucous, glabrous or pilose;

branchlets yellow-brown, gray-brown, or red-brown, densely short-silky, velvety, or villous, (buds caprea-type or intermediate).

branches red-brown, pubescent to glabrescent;

branchlets yellow-brown or red-brown, sparsely velvety or pubescent.

Leaves

stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 3–13(–16) mm, tomentose or velvety adaxially;

largest medial blade elliptic, narrowly oblanceolate, oblanceolate, or obovate, 31–70–120 × 17–48 mm, 2.1–3.1–4 times as long as wide, base cuneate or convex, margins slightly revolute or flat, strongly revolute proximally, entire, irregularly serrate, or sinuate, (glands submarginal or epilaminal), apex acuminate or convex, abaxial surface not evidently glaucous, (obscured by hairs), densely short-silky, woolly, or silky-woolly, hairs straight, wavy, or curved, adaxial slightly glossy (sometimes dull and glaucous), pilose or moderately densely short-silky;

proximal blade margins entire or shallowly serrulate;

juvenile blade green, densely long-silky or woolly abaxially, (sparsely silky-tomentose adaxially), hairs white.

stipules absent or rudimentary on early ones, foliaceous on late ones, apex rounded;

petiole (shallowly grooved adaxially), 13–20 mm, puberulent or velvety adaxially;

largest medial blade narrowly oblong, oblong, lanceolate, or narrowly ovate, 100–170 × 38–55 mm, 2.5–3(–4) times as long as wide, base rounded, convex, or subcordate (cordate on larger leaves), margins serrulate or spinulose-serrulate, apex acute, acuminate, or convex, abaxial surface sparsely tomentose, hairs straight, adaxial highly glossy, sparsely villous or pilose to glabrescent (midrib remaining villous), hairs white and ferruginous;

proximal blade margins entire;

juvenile blade sparsely pubescent to very densely villous or pilose abaxially, hairs white.

Catkins

flowering just before or as leaves emerge;

staminate slender or stout, (17–)22–54 × 8–15 mm, flowering branchlet 1–9 mm;

pistillate moderately densely flowered, slender to stout, 25–73(–115 in fruit) × 5–15 mm, flowering branchlet 1–20 mm;

floral bract tawny to dark brown, 1.4–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight or wavy.

staminate 29–72 × 12–15 mm, flowering branchlet 1–10 mm;

pistillate 50–81 × 17–27 mm, flowering branchlet 5–30 mm;

floral bract (tawny, sometimes greenish), 2–3.6 mm, apex rounded, entire, abaxially sparsely hairy, hairs wavy;

pistillate bract persistent after flowering.

Staminate flowers

adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.4–1.3 mm;

stamens 1;

filaments distinct, glabrous;

anthers purple turning yellow, shortly cylindrical, 0.5–0.7 mm.

abaxial nectary 0.4–0.8 mm, adaxial nectary ovate, 0.5–1.1 mm, nectaries distinct or connate and cup-shaped;

stamens 3–7;

filaments hairy basally;

anthers 0.4–0.5 mm.

Pistillate flowers

adaxial nectary square, ovate, or flask-shaped, 0.5–0.9 mm, shorter to longer than stipe;

stipe 0.4–1.4 mm;

ovary ovoid to pyriform, long- or short-silky or villous, beak sometimes slightly bulged below styles;

ovules 14–20 per ovary;

styles 0.4–0.8 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, 0.16–0.28–0.4 mm.

adaxial nectary square, 0.5–0.9 mm;

stipe 3.2–5.6 mm;

ovary obclavate to ellipsoidal, (rarely puberulent), beak gradually tapering to styles;

ovules 4 per ovary;

styles 0.3–0.4 mm;

stigmas 0.16–0.17–0.2 mm.

Capsules

3.5–5.6 mm.

6–7 mm.

2n

= 38.

= 38.

Salix sitchensis

Salix floridana

Phenology Flowering early Apr-mid Jun (Mar in California). Flowering mid Feb-early Apr.
Habitat Tidal swamps and marshes, coastal fog belts and headlands, sand dunes, springs, gravelly streambeds and deltas, glacial moraines, avalanche tracks, dry canyons, clearings and edges of forests, shade tolerant Swamps, marshy shores of streams in woodlands, calcareous areas, shade tolerant
Elevation 0-1800 m [0-5900 ft] 10-40 m [30-130 ft]
Distribution
from FNA
AK; CA; ID; MT; OR; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA
[BONAP county map]
Discussion

Ovary hairiness in some Salix sitchensis populations varies from uniformly hairy to glabrescent, with intermediates with patchy or streaky hairiness. All three variations can occur together and do not seem to indicate hybridization.

Both Salix sitchensis and S. scouleriana have similar variants with leaves having very densely curly hairs on abaxial surfaces [S. sitchensis forma coulteri (Andersson) Jepson and S. scouleriana forma poikila (C. K. Schneider) C. K. Schneider]. Plants resembling S. drummondiana but with similar indumentum probably are hybrids with S. alaxensis (see 84. S. drummondiana). The coulteri taxon resembles S. delnortensis in having stipules with adaxial surfaces glabrous and very sparsely glandular toward the base, densely hairy abaxially, and with gland-dotted margins; its branchlets have wavy to crinkly hairs. The possible hybrid origin of S. delnortensis needs study (R. D. Dorn 2000).

Hybrids:

Salix sitchensis forms natural hybrids with S. alaxensis var. longistylis and S. melanopsis. Hybridization with S. geyeriana reported by J. K. Henry (1915) is not based on convincing specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

The closest relatives of Salix floridana are in the Old World sect. Tetraspermae. A detailed discussion of the distribution, taxonomy, and relationships of this uncommon subtropical endemic was given by G. W. Argus (1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 159. Treatment author: George W. Argus. FNA vol. 7, p. 32. Treatment author: George W. Argus.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Sitchenses Salicaceae > Salix > subg. Protitea > sect. Floridanae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. coulteri, S. sitchensis var. parvifolia
Name authority Sanson ex Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 162. (1832) Chapman: Fl. South. U.S., 430. (1860)
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