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mountain willow, Scouler willow, Scouler's willow

bayberry willow, blue-leaf willow

Habit Shrubs or trees, 1–10(–20) m. Stems: branches gray-brown, yellow-brown, or red-brown, not glaucous, glabrous or tomentose; branchlets yellow-green or yellow-brown, sparsely to densely villous, tomentose, or velvety. Plants 0.3–5 m, (sometimes forming clones by stem fragmentation or layering).
Stems

branches (sometimes highly brittle at base), red-brown or yellow-brown, not or weakly glaucous, (slightly or highly glossy), glabrous or villous;

branchlets red-brown or yellow-brown, glabrous or sparsely to very densely villous, (buds caprea-type, inner membranaceous bud-scale layer free and separating or not).

Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones, (1–16 mm), apex acute or acuminate;

petiole convex to flat adaxially, 2–13 mm, velvety or villous adaxially;

largest medial blade usually oblanceolate, sometimes narrowly elliptic, elliptic or obovate, 29–100 × 9–37 mm, 1.7–3.9 times as long as wide, base cuneate or convex, margins strongly to slightly revolute or flat, entire, remotely serrate, crenate, or sinuate, (glands submarginal or epilaminal), apex acuminate, convex, or rounded, abaxial surface glaucous, sparsely to densely short- to long-silky or woolly, hairs (white, sometimes also ferruginous), wavy or straight, adaxial slightly glossy, pilose or moderately densely short-silky, midrib velutinous or villous, (hairs white, sometimes also ferruginous);

proximal blade margins entire, serrulate, or crenulate;

juvenile blade reddish or yellowish green, sparsely to densely villous, short- or long-silky abaxially, hairs white, sometimes also ferruginous.

stipules rudimentary or foliaceous on early ones, foliaceous on late ones, apex acute or acuminate;

petiole shallowly grooved, or convex to flat adaxially, 3.5–7.3–13 mm, (sometimes with 2 spherical glands distally), villous, tomentose, pilose, or pubescent adaxially;

largest medial blade narrowly oblong, narrowly elliptic, elliptic, or oblanceolate, 35–61.3–110 × 11–16–46 mm, 2–2.7–5.2 times as long as wide, base convex, rounded, subcordate, or cuneate, margins flat or slightly revolute, (thickened and raised), crenulate or serrulate, apex acuminate, acute, or convex, abaxial surface usually very thickly glaucous, glabrous or pilose, midribs pubescent to tomentose, hairs (white, often also ferruginous), curved, wavy, or straight, adaxial slightly glossy, glabrous or pilose, midribs sparsely pubescent (hairs white, sometimes also ferruginous);

proximal blade margins entire or serrulate;

juvenile blade translucent, reddish or yellowish green, glabrous or sparsely pubescent abaxially, midribs often densely hairy, hairs white, sometimes also ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.4–0.9 mm;

filaments distinct, glabrous or hairy on proximal 1/2;

anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.7–1.2 mm.

adaxial nectary narrowly oblong, oblong, square, or ovate, 0.44–0.48–1.4 mm;

filaments distinct, glabrous;

anthers yellow, (ellipsoid or globose), 0.52–0.69–0.76 mm.

Pistillate flowers

adaxial nectary oblong or square, 0.2–0.8 mm, shorter than stipe;

stipe 0.8–2.3 mm;

ovary pyriform or obclavate, densely long-silky, beak slightly bulged below styles;

ovules 10–18 per ovary;

styles 0.2–0.6 mm;

stigmas slenderly cylindrical, 0.4–0.82–1.04 mm.

adaxial nectary oblong, narrowly oblong, square, or flask-shaped, 0.56–0.8–1.4 mm, shorter than stipe;

stipe 0.96–1.7–3.4 mm;

ovary pyriform, glabrous, beak slightly bulged below styles;

ovules 12–14 per ovary;

styles (sometimes distinct), 0.3–0.8–1.3 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or slenderly cylindrical, 0.24–0.43–0.56 mm.

Capsules

4.5–11 mm.

5–7–11 mm.

Catkins

flowering before leaves emerge; staminate stout or subglobose, 18–40.5 × 8–22 mm, flowering branchlet 0–4 mm; pistillate very densely flowered, slender or stout, 18–60(–90 in fruit) × 10–22 mm, flowering branchlet 0–8 mm;

floral bract brown, black, or bicolor, 1.5–4.5 mm, apex rounded or acute, abaxially hairy, hairs straight.

staminate flowering before leaves emerge, pistillate as leaves emerge; staminate stout or slender, 23.5–35.6–51 × 9–12.7–22 mm, flowering branchlet 1–3.9–10 mm; pistillate loosely flowered, stout or slender, 19–42–62(–85 in fruit) × 8–13–18 mm, flowering branchlet 1.5–5.9–13 mm;

floral bract brown or bicolor, 1.2–1.8–3 mm, apex rounded or acute, sometimes toothed, abaxially hairy, hairs straight or wavy.

2n

= 76.

Salix scouleriana

Salix myricoides

Phenology Flowering late Feb-mid Jun. Flowering early Apr-early Jul.
Habitat Dry conifer forests, mature woods on edges of streams and lakes, treed bogs, meadows, subalpine slopes, springs, pine barrens, openings in old burns, arroyos and disturbed sites, sandy, silty-clay, or gravelly, igneous substrates Stream and lake shores, gravel bars, subalpine conifer forests, alkaline fens, sea cliffs, dry limestone talus, swamps, tidal meadows, sand dunes
Elevation 0-3500 m (0-11500 ft) 0-1100 m (0-3600 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
IL; IN; ME; MI; OH; PA; WI; NB; NF; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Western Salix scouleriana and eastern S. humilis are closely related and are sometimes difficult to separate. Although there is an apparent range disjunction between them in western Manitoba, it may be a collecting gap. In general, S. scouleriana differs from S. humilis in being a taller shrub, sometimes even tree-like, with broader leaves and longer catkins, floral bracts, stigmas, and styles, but these quantitative characteristics all overlap. The apparent difference in anther length (S. scouleriana 0.7–1.2 mm; S. humilis 0.4–0.6 mm) may be correlated with a difference in chromosome number. Salix scouleriana is tetraploid (Y. Suda and G. W. Argus 1968); S. humilis has been reported to be both diploid (Suda and Argus; L. Zsuffa and Y. Raj, unpubl.) and tetraploid (R. D. Dorn 1976). The latter count was from the same population as the one by Suda and Argus. Further chromosome counts are indicated.

See 77. Salix hookeriana for comparative descriptions.

Hybrids:

Salix scouleriana forms natural hybrids with S. hookeriana, S. planifolia, and S. pulchra.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants with densely villous branchlets and branches have been named var. albovestita. Branchlet indumentum varies widely in the species and seems to be continuous, with both villous and glabrous variants sometimes occurring in the same area. This characteristic is more common in populations on the shores of the Great Lakes and on the western coast of James Bay, but even these populations are variable.

Reports of Salix myricoides from Akimiski Island, Nunavut, were based on misidentified S. planifolia. See 68. S. eriocephala for differences.

Hybrids:

Salix myricoides forms natural hybrids with S. bebbiana, S. discolor, and S. glauca var. cordifolia. Hybrids with S. eriocephala have been reported (M. L. Fernald 1950) but no convincing specimens have been seen. Inasmuch as S. eriocephala and S. myricoides are very similar, hybrids between the two would be very difficult to identify.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 131. FNA vol. 7, p. 119.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Cinerella Salicaceae > Salix > subg. Vetrix > sect. Hastatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. scouleriana var. poikila S. glaucophylla var. albovestita, S. glaucophylloides, S. glaucophylloides var. albovestita, S. glaucophylloides var. glaucophylla, S. myricoides var. albovestita
Name authority Barratt ex Hooker: Fl. Bor.-Amer. 2: 145. (1838) Muhlenberg: Ges. Naturf. Freunde Berlin Neue Schriften 4: 235, plate 6, fig. 2. (1803)
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