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mountain willow, Scouler willow, Scouler's willow

Lemmon's willow

Habit Shrubs or trees, 1–10(–20) m. Stems: branches gray-brown, yellow-brown, or red-brown, not glaucous, glabrous or tomentose; branchlets yellow-green or yellow-brown, sparsely to densely villous, tomentose, or velvety. Plants 1.2–4 m, (sometimes forming clones by stem fragmentation).
Stems

branches (sometimes highly brittle at base), yellow-brown or red-brown, not to sometimes strongly glaucous, (dull to slightly glossy), glabrous or puberulent at nodes;

branchlets yellow-brown to red-brown, (weakly or strongly glaucous or not), puberulent, (buds alba-type or intermediate, scale inner membranaceous layer free, not separating from outer layer).

Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones, (1–16 mm), apex acute or acuminate;

petiole convex to flat adaxially, 2–13 mm, velvety or villous adaxially;

largest medial blade usually oblanceolate, sometimes narrowly elliptic, elliptic or obovate, 29–100 × 9–37 mm, 1.7–3.9 times as long as wide, base cuneate or convex, margins strongly to slightly revolute or flat, entire, remotely serrate, crenate, or sinuate, (glands submarginal or epilaminal), apex acuminate, convex, or rounded, abaxial surface glaucous, sparsely to densely short- to long-silky or woolly, hairs (white, sometimes also ferruginous), wavy or straight, adaxial slightly glossy, pilose or moderately densely short-silky, midrib velutinous or villous, (hairs white, sometimes also ferruginous);

proximal blade margins entire, serrulate, or crenulate;

juvenile blade reddish or yellowish green, sparsely to densely villous, short- or long-silky abaxially, hairs white, sometimes also ferruginous.

stipules absent or rudimentary on early ones, foliaceous on late ones, apex acute;

petiole convex to flat adaxially, 5–16 mm, tomentose or velvety adaxially;

largest medial blade (sometimes amphistomatous), lorate or very narrowly to narrowly elliptic, 44–110 × 6–22 mm, 3.4–9.9(–12) times as long as wide, base convex or cuneate, margins flat to slightly revolute, entire or shallowly serrulate, apex acuminate to acute, abaxial surface glaucous, sparsely short- or long-silky to glabrescent, hairs (white, also ferruginous), straight or wavy, adaxial slightly glossy, sparsely short-silky, (hairs also ferruginous);

proximal blade margins entire or shallowly serrulate;

juvenile blade reddish or yellowish green, densely long-silky abaxially, hairs white and ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.4–0.9 mm;

filaments distinct, glabrous or hairy on proximal 1/2;

anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.7–1.2 mm.

adaxial nectary narrowly oblong, oblong, or ovate, 0.3–0.9 mm;

filaments distinct or connate less than 1/2 their lengths, glabrous or hairy on proximal 1/2 or basally;

anthers yellow, ellipsoid, shortly cylindrical, or globose, 0.5–0.9 mm.

Pistillate flowers

adaxial nectary oblong or square, 0.2–0.8 mm, shorter than stipe;

stipe 0.8–2.3 mm;

ovary pyriform or obclavate, densely long-silky, beak slightly bulged below styles;

ovules 10–18 per ovary;

styles 0.2–0.6 mm;

stigmas slenderly cylindrical, 0.4–0.82–1.04 mm.

adaxial nectary oblong, narrowly oblong, or ovate, 0.4–1 mm;

stipe 1.1–2.1 mm;

ovary pyriform, beak sometimes slightly bulged below styles;

ovules 12 per ovary;

styles 0.3–1 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.2–0.3–0.48 mm.

Capsules

4.5–11 mm.

5–7 mm.

Catkins

flowering before leaves emerge; staminate stout or subglobose, 18–40.5 × 8–22 mm, flowering branchlet 0–4 mm; pistillate very densely flowered, slender or stout, 18–60(–90 in fruit) × 10–22 mm, flowering branchlet 0–8 mm;

floral bract brown, black, or bicolor, 1.5–4.5 mm, apex rounded or acute, abaxially hairy, hairs straight.

flowering just before or, sometimes, as leaves emerge; staminate stout, (1.4–2.7–)16–28 × 9–17 mm, flowering branchlet 1–3 mm; pistillate loosely flowered, stout, (1.5–2.7–)19–44(–65 in fruit) × 10–18 mm, flowering branchlet 1–6 mm;

floral bract brown or bicolor, 1.1–2.4 mm, apex rounded or convex, abaxially hairy, hairs straight.

2n

= 76.

= 76.

Salix scouleriana

Salix lemmonii

Phenology Flowering late Feb-mid Jun. Flowering early Mar-late Jun.
Habitat Dry conifer forests, mature woods on edges of streams and lakes, treed bogs, meadows, subalpine slopes, springs, pine barrens, openings in old burns, arroyos and disturbed sites, sandy, silty-clay, or gravelly, igneous substrates Streams, lakeshores, wet meadows, springs, old burns in subalpine conifer forests, sandy, granite substrates
Elevation 0-3500 m (0-11500 ft) 1400-3500 m (4600-11500 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; NV; OR; WY; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Western Salix scouleriana and eastern S. humilis are closely related and are sometimes difficult to separate. Although there is an apparent range disjunction between them in western Manitoba, it may be a collecting gap. In general, S. scouleriana differs from S. humilis in being a taller shrub, sometimes even tree-like, with broader leaves and longer catkins, floral bracts, stigmas, and styles, but these quantitative characteristics all overlap. The apparent difference in anther length (S. scouleriana 0.7–1.2 mm; S. humilis 0.4–0.6 mm) may be correlated with a difference in chromosome number. Salix scouleriana is tetraploid (Y. Suda and G. W. Argus 1968); S. humilis has been reported to be both diploid (Suda and Argus; L. Zsuffa and Y. Raj, unpubl.) and tetraploid (R. D. Dorn 1976). The latter count was from the same population as the one by Suda and Argus. Further chromosome counts are indicated.

See 77. Salix hookeriana for comparative descriptions.

Hybrids:

Salix scouleriana forms natural hybrids with S. hookeriana, S. planifolia, and S. pulchra.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

See 85. Salix drummondiana for comments on differences.

Hybrids:

Salix lemmonii forms natural hybrids with S. geyeriana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 131. FNA vol. 7, p. 154.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Cinerella Salicaceae > Salix > subg. Vetrix > sect. Geyerianae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. scouleriana var. poikila
Name authority Barratt ex Hooker: Fl. Bor.-Amer. 2: 145. (1838) Bebb: Willows Calif., 88. (1879)
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