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mountain willow, Scouler willow, Scouler's willow

Goodding's black willow, Goodding's willow, Gooding's willow

Habit Shrubs or trees, 1–10(–20) m. Stems: branches gray-brown, yellow-brown, or red-brown, not glaucous, glabrous or tomentose; branchlets yellow-green or yellow-brown, sparsely to densely villous, tomentose, or velvety. Trees, 3–30 m. Stems: branches flexible to ± brittle at base, yellow-brown to gray-brown, pubescent to glabrescent; branchlets usually yellowish or yellow-green, sometimes reddish brown, puberulent or pubescent to glabrescent.
Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones, (1–16 mm), apex acute or acuminate;

petiole convex to flat adaxially, 2–13 mm, velvety or villous adaxially;

largest medial blade usually oblanceolate, sometimes narrowly elliptic, elliptic or obovate, 29–100 × 9–37 mm, 1.7–3.9 times as long as wide, base cuneate or convex, margins strongly to slightly revolute or flat, entire, remotely serrate, crenate, or sinuate, (glands submarginal or epilaminal), apex acuminate, convex, or rounded, abaxial surface glaucous, sparsely to densely short- to long-silky or woolly, hairs (white, sometimes also ferruginous), wavy or straight, adaxial slightly glossy, pilose or moderately densely short-silky, midrib velutinous or villous, (hairs white, sometimes also ferruginous);

proximal blade margins entire, serrulate, or crenulate;

juvenile blade reddish or yellowish green, sparsely to densely villous, short- or long-silky abaxially, hairs white, sometimes also ferruginous.

stipules broad rudiments or foliaceous on early ones, foliaceous on late ones, (glands numerous adaxially), apex rounded to convex;

petiole (sometimes with spherical glands distally), 4–10 mm, pilose adaxially;

largest medial blade (sometimes amphistomatous), narrowly elliptic, very broadly oblong, lorate, or linear, 67–130 × 9.5–16 mm, 4.7–12.4 times as long as wide, base cuneate to convex, margins serrulate to serrate, apex acuminate, caudate, or acute, abaxial surface (usually not glaucous, rarely thinly so), glabrous or puberulent, hairs wavy, adaxial slightly glossy, pilose to glabrescent;

proximal blade margins entire or shallowly serrulate;

juvenile blade sparsely velvety to pilose abaxially, hairs white.

Staminate flowers

adaxial nectary oblong or square, 0.4–0.9 mm;

filaments distinct, glabrous or hairy on proximal 1/2;

anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.7–1.2 mm.

abaxial nectary (0.2–)0.3–0.6 mm, adaxial nectary square to ovate, 0.2–0.6 mm, nectaries distinct;

stamens 4–6(–8);

filaments (sometimes basally connate), hairy on proximal 1/2;

anthers 0.4–0.5 mm, (axes straight).

Pistillate flowers

adaxial nectary oblong or square, 0.2–0.8 mm, shorter than stipe;

stipe 0.8–2.3 mm;

ovary pyriform or obclavate, densely long-silky, beak slightly bulged below styles;

ovules 10–18 per ovary;

styles 0.2–0.6 mm;

stigmas slenderly cylindrical, 0.4–0.82–1.04 mm.

adaxial nectary square (flattened), 0.2–0.6 mm;

stipe 1.2–3.2 mm;

ovary pyriform, (sometimes villous), beak slightly bulged or abruptly tapering to styles;

ovules 12–18 per ovary;

styles 0.1–0.3 mm;

stigmas 0.2–0.29–0.32 mm.

Capsules

4.5–11 mm.

6–7 mm.

Catkins

flowering before leaves emerge; staminate stout or subglobose, 18–40.5 × 8–22 mm, flowering branchlet 0–4 mm; pistillate very densely flowered, slender or stout, 18–60(–90 in fruit) × 10–22 mm, flowering branchlet 0–8 mm;

floral bract brown, black, or bicolor, 1.5–4.5 mm, apex rounded or acute, abaxially hairy, hairs straight.

staminate 19–80 × 6–10 mm, flowering branchlet 2–23 mm; pistillate 23–82 × 6–15 mm, flowering branchlet 2–48 mm;

floral bract 1.4–2.4 mm, apex acute or rounded, entire or toothed, abaxially sparsely to moderately densely hairy, hairs wavy; pistillate bract deciduous after flowering.

2n

= 76.

= 38.

Salix scouleriana

Salix gooddingii

Phenology Flowering late Feb-mid Jun. Flowering late Mar–Jun.
Habitat Dry conifer forests, mature woods on edges of streams and lakes, treed bogs, meadows, subalpine slopes, springs, pine barrens, openings in old burns, arroyos and disturbed sites, sandy, silty-clay, or gravelly, igneous substrates Riparian forests, springs, seepage areas, washes, meadows
Elevation 0-3500 m (0-11500 ft) -40-500(-2500) m (-100-1600(-8200) ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NM; NV; OK; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Guerrero, Sinaloa, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Western Salix scouleriana and eastern S. humilis are closely related and are sometimes difficult to separate. Although there is an apparent range disjunction between them in western Manitoba, it may be a collecting gap. In general, S. scouleriana differs from S. humilis in being a taller shrub, sometimes even tree-like, with broader leaves and longer catkins, floral bracts, stigmas, and styles, but these quantitative characteristics all overlap. The apparent difference in anther length (S. scouleriana 0.7–1.2 mm; S. humilis 0.4–0.6 mm) may be correlated with a difference in chromosome number. Salix scouleriana is tetraploid (Y. Suda and G. W. Argus 1968); S. humilis has been reported to be both diploid (Suda and Argus; L. Zsuffa and Y. Raj, unpubl.) and tetraploid (R. D. Dorn 1976). The latter count was from the same population as the one by Suda and Argus. Further chromosome counts are indicated.

See 77. Salix hookeriana for comparative descriptions.

Hybrids:

Salix scouleriana forms natural hybrids with S. hookeriana, S. planifolia, and S. pulchra.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix gooddingii and S. nigra are closely related and are sometimes treated as conspecific (C. R. Ball 1950). Salix gooddingii has yellow-brown or pale gray-brown branches, capsules 6–7 mm, and ovaries usually glabrous but pilose in ca. 20% of specimens. Salix nigra has red-brown to dark gray-brown branches, capsules 3–5 mm, and ovaries almost always glabrous. A single plant with pilose ovaries was found in Ontario, Canada; reports (W. A. Archer 1965) of S. nigra with hairy ovaries in Alabama, Arkansas, Illinois, Iowa, Kansas, Massachusetts, Oklahoma, and Texas could not be confirmed. Ranges of these taxa overlap in west-central Texas, where there is evidence of intergradation; they rarely occur in the same population. The map by E. L. Little Jr. (1971), who treated them as conspecific, shows a significant range disjunction between the two.

Catkins of Salix gooddingii flowering in March and early April are sometimes borne in leaf axils. This suggests that the sylleptic condition, typical of S. bonplandiana, is sometimes ecotypic.

Hybrids:

Salix gooddingii forms natural hybrids with S. amygdaloides and S. nigra. Hybrids with S. lasiandra have been reported (C. K. Schneider 1921); no convincing specimens have been seen.

Salix gooddingii × S. laevigata: In Arizona, a population of young plants displays intermediate characteristics. They have leaf blades sparsely glaucous abaxially, as in S. laevigata, but narrow, often amphistomatous, and with petioles sometimes not glandular distally, as in S. gooddingii. Both parental species occur in the region. This hybrid was also reported by C. K. Schneider (1921) from California.

Salix gooddingii × S. nigra: This hybrid may occur in western Texas where the parental species overlap. Some specimens from that area seem to be “intermediate” in branch color, but the differences are subtle.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 131. FNA vol. 7, p. 36.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Cinerella Salicaceae > Salix > subg. Protitea > sect. Humboldtianae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. scouleriana var. poikila S. gooddingii var. vallicola, S. gooddingii var. variabilis, S. nigra var. vallicola
Name authority Barratt ex Hooker: Fl. Bor.-Amer. 2: 145. (1838) C. R. Ball: Bot. Gaz. 40: 376, plate 12, figs. 1, 2. (1905)
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