Salix scouleriana |
Salix glauca |
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mountain willow, Scouler willow, Scouler's willow |
glaucous willow, gray willow, gray-leaf willow, grey-leaf willow |
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Habit | Shrubs or trees, 1–10(–20) m. Stems: branches gray-brown, yellow-brown, or red-brown, not glaucous, glabrous or tomentose; branchlets yellow-green or yellow-brown, sparsely to densely villous, tomentose, or velvety. | Plants 0.2–6 m, not clonal. | ||||||||||||
Stems | erect or decumbent; branches brownish, yellow-brown, gray-brown, or red-brown, villous or pilose to glabrescent; branchlets yellow-brown or red-brown, sparsely to densely villous or tomentose to glabrescent. |
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Leaves | stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones, (1–16 mm), apex acute or acuminate; petiole convex to flat adaxially, 2–13 mm, velvety or villous adaxially; largest medial blade usually oblanceolate, sometimes narrowly elliptic, elliptic or obovate, 29–100 × 9–37 mm, 1.7–3.9 times as long as wide, base cuneate or convex, margins strongly to slightly revolute or flat, entire, remotely serrate, crenate, or sinuate, (glands submarginal or epilaminal), apex acuminate, convex, or rounded, abaxial surface glaucous, sparsely to densely short- to long-silky or woolly, hairs (white, sometimes also ferruginous), wavy or straight, adaxial slightly glossy, pilose or moderately densely short-silky, midrib velutinous or villous, (hairs white, sometimes also ferruginous); proximal blade margins entire, serrulate, or crenulate; juvenile blade reddish or yellowish green, sparsely to densely villous, short- or long-silky abaxially, hairs white, sometimes also ferruginous. |
stipules (marcescent or not), foliaceous or rudimentary on early and late ones; petiole (usually deeply to shallowly grooved adaxially), 1–27 mm, (much longer than subtended bud); largest medial blade usually hypostomatous, sometimes hemiamphistomatous or amphistomatous, usually narrowly elliptic, elliptic, usually oblanceolate or obovate, sometimes narrowly oblong or obovate, 27–82 × 6–39 mm, 1.4–4.8 times as long as wide, base usually cuneate or convex, sometimes rounded, rarely subcordate, margins slightly revolute or flat, usually entire, apex acute, acuminate, convex, or rounded, abaxial surface densely villous or villous-silky, tomentose, short- or long-silky, or pilose, hairs usually wavy or straight, sometimes curved, adaxial usually slightly glossy, sometimes dull, moderately densely villous, pilose, or long-silky to glabrescent; proximal blade margins entire or serrulate; juvenile blade sparsely or densely villous, tomentose, or long-silky abaxially. |
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Staminate flowers | adaxial nectary oblong or square, 0.4–0.9 mm; filaments distinct, glabrous or hairy on proximal 1/2; anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.7–1.2 mm. |
abaxial nectary 0.1–1 mm, adaxial nectary narrowly oblong, oblong, square, ovate, or flask-shaped, 0.5–1.3 mm, nectaries distinct, or connate and cup-shaped; filaments distinct or slightly or partly connate, glabrous, or hairy on proximal 1/2; anthers 0.4–0.8 mm. |
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Pistillate flowers | adaxial nectary oblong or square, 0.2–0.8 mm, shorter than stipe; stipe 0.8–2.3 mm; ovary pyriform or obclavate, densely long-silky, beak slightly bulged below styles; ovules 10–18 per ovary; styles 0.2–0.6 mm; stigmas slenderly cylindrical, 0.4–0.82–1.04 mm. |
abaxial nectary absent, adaxial nectary usually narrowly oblong, oblong, or ovate, sometimes flask-shaped, 0.4–1.8 mm, shorter to longer than stipe; stipe 0.3–2.8 mm; ovary pyriform or obclavate, densely villous, tomentose, short-silky, or pubescent, beak usually gradually tapering to styles, sometimes gradually tapering to or slightly bulged below styles; ovules 6–22 per ovary; styles connate to distinct 1/2 their lengths or more, 0.3–1.6 mm; stigmas flat, abaxially non-papillate with rounded tip, or slenderly or broadly cylindrical, 0.2–0.8 mm. |
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Capsules | 4.5–11 mm. |
4.5–9 mm. |
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Catkins | flowering before leaves emerge; staminate stout or subglobose, 18–40.5 × 8–22 mm, flowering branchlet 0–4 mm; pistillate very densely flowered, slender or stout, 18–60(–90 in fruit) × 10–22 mm, flowering branchlet 0–8 mm; floral bract brown, black, or bicolor, 1.5–4.5 mm, apex rounded or acute, abaxially hairy, hairs straight. |
staminate 14–53 × 5–17 mm, flowering branchlet 1–25 mm; pistillate densely to sometimes loosely flowered, slender, stout, subglobose, or globose, 15–83 × 7–21 mm, flowering branchlet 2–37 mm; floral bract tawny, brown, bicolor, or greenish, 1–3.4 mm, apex convex or rounded, entire, abaxially hairy, hairs wavy, crinkled, or straight. |
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2n | = 76. |
= 76, 95, 114, 152. |
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Salix scouleriana |
Salix glauca |
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Phenology | Flowering late Feb-mid Jun. | |||||||||||||
Habitat | Dry conifer forests, mature woods on edges of streams and lakes, treed bogs, meadows, subalpine slopes, springs, pine barrens, openings in old burns, arroyos and disturbed sites, sandy, silty-clay, or gravelly, igneous substrates | |||||||||||||
Elevation | 0-3500 m (0-11500 ft) | |||||||||||||
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Mexico (Chihuahua, Sonora)
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AK; CO; MT; NM; OR; UT; WA; WY; AB; BC; MB; NS; NT; NU; ON; QC; SK; YT; Eurasia (China [Altay Shan], Chukotka, Mongolia, Novaya Zemlya, Russian Far East, Sakhalin, Scandinavia, arctic, e, w Siberia)
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Discussion | Western Salix scouleriana and eastern S. humilis are closely related and are sometimes difficult to separate. Although there is an apparent range disjunction between them in western Manitoba, it may be a collecting gap. In general, S. scouleriana differs from S. humilis in being a taller shrub, sometimes even tree-like, with broader leaves and longer catkins, floral bracts, stigmas, and styles, but these quantitative characteristics all overlap. The apparent difference in anther length (S. scouleriana 0.7–1.2 mm; S. humilis 0.4–0.6 mm) may be correlated with a difference in chromosome number. Salix scouleriana is tetraploid (Y. Suda and G. W. Argus 1968); S. humilis has been reported to be both diploid (Suda and Argus; L. Zsuffa and Y. Raj, unpubl.) and tetraploid (R. D. Dorn 1976). The latter count was from the same population as the one by Suda and Argus. Further chromosome counts are indicated. See 77. Salix hookeriana for comparative descriptions. Hybrids: Salix scouleriana forms natural hybrids with S. hookeriana, S. planifolia, and S. pulchra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 5 (4 in the flora). Occurrence of Salix glauca is disjunct in western Siberia; var. glauca is known from Scandinavia. Salix glauca is ubiquitous, highly polymorphic, and polyploid. It appears to have four major variations that are treated in different ways. European floras have treated it as an undivided polymorphic species(A. K. Skvortsov 1999), as several distinct species (K. H. Rechinger 1964b), or as comprising several subspecies (Rechinger 1993; G. W. Argus et al. 1999; B. Jonsell and T. Karlsson 2000+, vol. 1). In North America, Argus (1965) treated the four central tendencies as widely intergrading “phases” of a single species, later adopting varietal rank, whereas E. Hultén (1968) recognized them as subspecies. In view of the fact that the four elements are confluent over wide areas, there may be some merit in recognizing them as informal phases, but varietal rank is used here. All specimens from Iceland named Salix glauca are S. arctica and those from Svalbard, Norway, are S. lanata Linnaeus. The major reason for the high variability within Salix glauca seems to be high, and probably recurrent, polyploidy. Tetraploids, pentaploids, hexaploids, and octoploids are known in the species, and two of the subspecies include three ploidal levels. There also are intergrading geographical variations that are the basis for the recognition of infraspecific taxa. Hybrids: Salix glauca forms natural hybrids with S. arctica, S. arctophila, S. ballii, S. barclayi, S. boothii, S. brachycarpa, S. eastwoodiae, S. myricoides, S. niphoclada, S. pedicellaris, and S. planifolia. The following key will help identify the varieties, but there are extensive areas of overlap among them and many intermediates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 131. | FNA vol. 7, p. 89. | ||||||||||||
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Cinerella | Salicaceae > Salix > subg. Chamaetia > sect. Glaucae | ||||||||||||
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Synonyms | S. scouleriana var. poikila | |||||||||||||
Name authority | Barratt ex Hooker: Fl. Bor.-Amer. 2: 145. (1838) | Linnaeus: Sp. Pl. 2: 1019. (1753) | ||||||||||||
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