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mountain willow, Scouler willow, Scouler's willow

Columbia River willow

Habit Shrubs or trees, 1–10(–20) m. Stems: branches gray-brown, yellow-brown, or red-brown, not glaucous, glabrous or tomentose; branchlets yellow-green or yellow-brown, sparsely to densely villous, tomentose, or velvety. Shrubs, 2–6.5 m. Stems: branches red-brown, glabrous or hairy at nodes; branchlets yellow-brown or red-brown, pubescent, puberulent, or glabrous, (hairs appressed, straight or wavy).
Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones, (1–16 mm), apex acute or acuminate;

petiole convex to flat adaxially, 2–13 mm, velvety or villous adaxially;

largest medial blade usually oblanceolate, sometimes narrowly elliptic, elliptic or obovate, 29–100 × 9–37 mm, 1.7–3.9 times as long as wide, base cuneate or convex, margins strongly to slightly revolute or flat, entire, remotely serrate, crenate, or sinuate, (glands submarginal or epilaminal), apex acuminate, convex, or rounded, abaxial surface glaucous, sparsely to densely short- to long-silky or woolly, hairs (white, sometimes also ferruginous), wavy or straight, adaxial slightly glossy, pilose or moderately densely short-silky, midrib velutinous or villous, (hairs white, sometimes also ferruginous);

proximal blade margins entire, serrulate, or crenulate;

juvenile blade reddish or yellowish green, sparsely to densely villous, short- or long-silky abaxially, hairs white, sometimes also ferruginous.

stipules usually absent or rudimentary, sometimes foliaceous on early ones (apex acute), rudimentary or foliaceous on late ones;

petiole 2–5 mm, short-silky adaxially;

largest medial blade (sometimes hypostomatous, glands marginal), linear or very narrowly elliptic, 58–115 × 5–17 mm, 5.7–12.8 times as long as wide, base acute or cuneate, margins flat to slightly revolute, remotely spinulose-serrulate or sinuate, apex acuminate, abaxial surface glaucous (sometimes obscured by hairs), sparsely to very densely short- or long-silky, hairs appressed to slightly spreading, straight, adaxial dull, pubescent or long-silky;

proximal blade margins entire or remotely denticulate;

juvenile blade color obscured by hairs, densely short-silky abaxially.

Staminate flowers

adaxial nectary oblong or square, 0.4–0.9 mm;

filaments distinct, glabrous or hairy on proximal 1/2;

anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.7–1.2 mm.

abaxial nectary 0.3–0.9 mm, adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.5–1 mm, nectaries distinct or connate and shallowly cup-shaped;

filaments hairy on proximal 1/2, throughout, or basally;

anthers 0.8–1.3 mm.

Pistillate flowers

adaxial nectary oblong or square, 0.2–0.8 mm, shorter than stipe;

stipe 0.8–2.3 mm;

ovary pyriform or obclavate, densely long-silky, beak slightly bulged below styles;

ovules 10–18 per ovary;

styles 0.2–0.6 mm;

stigmas slenderly cylindrical, 0.4–0.82–1.04 mm.

(abaxial nectary 0.3–0.4 mm), adaxial nectary narrowly oblong, ovate, or flask-shaped, 0.4–1.3 mm, longer than stipe, nectaries distinct or connate and shallowly or partially cup-shaped;

stipe 0.2–0.7 mm;

ovary pyriform, long-silky, beak abruptly tapering to styles;

ovules 18–30 per ovary;

styles 0.1–0.2(–0.4) mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, (0.3–)0.5–1.1 mm.

Capsules

4.5–11 mm.

3.4–5.6 mm.

Catkins

flowering before leaves emerge; staminate stout or subglobose, 18–40.5 × 8–22 mm, flowering branchlet 0–4 mm; pistillate very densely flowered, slender or stout, 18–60(–90 in fruit) × 10–22 mm, flowering branchlet 0–8 mm;

floral bract brown, black, or bicolor, 1.5–4.5 mm, apex rounded or acute, abaxially hairy, hairs straight.

staminate 20–83 × 5–13 mm, flowering branchlet 6–125(–200) mm; pistillate densely to moderately densely flowered, slender or stout, 35–90 × 5–12 mm, flowering branchlet 11–160 mm;

floral bract 1.4–4 mm, apex acute, rounded, retuse, or truncate, entire or toothed, abaxially hairy throughout or proximally, hairs straight or wavy.

2n

= 76.

= 38.

Salix scouleriana

Salix columbiana

Phenology Flowering late Feb-mid Jun. Flowering May-late Jul.
Habitat Dry conifer forests, mature woods on edges of streams and lakes, treed bogs, meadows, subalpine slopes, springs, pine barrens, openings in old burns, arroyos and disturbed sites, sandy, silty-clay, or gravelly, igneous substrates Riparian, sandy-silty to sandy gravel floodplains, old beach dunes, rocky fill along streams
Elevation 0-3500 m (0-11500 ft) 5-40 m (0-100 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; WA
[BONAP county map]
Discussion

Western Salix scouleriana and eastern S. humilis are closely related and are sometimes difficult to separate. Although there is an apparent range disjunction between them in western Manitoba, it may be a collecting gap. In general, S. scouleriana differs from S. humilis in being a taller shrub, sometimes even tree-like, with broader leaves and longer catkins, floral bracts, stigmas, and styles, but these quantitative characteristics all overlap. The apparent difference in anther length (S. scouleriana 0.7–1.2 mm; S. humilis 0.4–0.6 mm) may be correlated with a difference in chromosome number. Salix scouleriana is tetraploid (Y. Suda and G. W. Argus 1968); S. humilis has been reported to be both diploid (Suda and Argus; L. Zsuffa and Y. Raj, unpubl.) and tetraploid (R. D. Dorn 1976). The latter count was from the same population as the one by Suda and Argus. Further chromosome counts are indicated.

See 77. Salix hookeriana for comparative descriptions.

Hybrids:

Salix scouleriana forms natural hybrids with S. hookeriana, S. planifolia, and S. pulchra.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybrids:

Salix columbiana forms natural hybrids with S. exigua var. exigua and S. sessilifolia. Both hybrids are reported from Oregon and Washington (R. D. Dorn 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 131. FNA vol. 7, p. 59.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Cinerella Salicaceae > Salix > subg. Longifoliae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. scouleriana var. poikila S. exigua var. columbiana
Name authority Barratt ex Hooker: Fl. Bor.-Amer. 2: 145. (1838) (Dorn) Argus: Harvard Pap. Bot. 12: 359. (2007)
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